PEREZ FARFANTK: AMERICAN SOLENOCERID SHRIMPS 



in the synonymy of//, laeuis. In this same contri- 

 bution, Burkenroad mentioned a "minute denticle 

 of variable size" posterior to the epigastric tooth, 

 and suggested that it is "probably the remains of 

 the larval anterior dorsal organ." In that location, 

 however, I have observed nothing more than an 

 extremely slight elevation of the postrostral 

 carina, and that only in two specimens. Had this 

 feature not been mentioned by Burkenroad, I 

 should have overlooked it, and, after observing it, 

 I believe it to be insignificant. 



Bouvier (1906b) described Haliporus andro- 

 gynus on the basis of two specimens which bear, in 

 addition to a fully developed thelycum, both 

 petasma and appendices masculinae. Burkenroad 

 (1936) stated that the simultaneous presence of 

 the female and male external genitalia in these 

 specimens probably represents an abnormality. 

 Recently, Crosnier and Forest (1973) indicated 

 that this combination of secondary sexual charac- 

 ters could represent an expression of protandric 

 hermaphroditism, as reported by Heegaard ( 1967 ) 

 in Solenocera membranacea (Risso 1816). They 

 added that in a rather large number of penaeids 

 the maximum size of males corresponds to the 

 minimum size of females. It should be noted, how- 

 ever, that Burkenroad (1936) cited a female of 

 H. laevis, also examined by me, with a carapace 

 length of 8 mm, which is about half the length of 

 the largest known male, 15 mm cl. 



In the two syntypes of H. androgynus, which 

 have a carapace length of about 20 mm, the petas- 

 mata are shorter than in other males of equal 

 size, and exhibit and armature with these juvenile 

 features. In one of the specimens, the mesial pro- 

 jection is extremely small, whereas the lateral 

 projection and the distolateral process are well 

 developed; in the other, the mesial projection is 

 distinct, the lateral one very small, the disto- 



lateral process is well developed, and a subdistal 

 spine is present on the free margin of the ventral 

 costa. 



Hymenopenaeus doris (Faxon 1893) 



Figures 9, 17, ISA, 19-20 



Haliporus doris Faxon 1893:214 [syntypes: 4 2, 

 MCZ 4648, off Cabo Velas Costa Rica, 10°14'N, 

 96°28'W, 2,232 fm (4,082 m), 8 April 1891, 

 Albatross stn 3414. 1 2, USNM 21182, S of 

 Punta Maldonado, Guerrero, Mexico, 14°46'N, 

 98°40'W, 1,879 fm (3,437 m), 10 April 1891, 

 Albatross stn 3415]. Faxon 1895:191, pi. 49, 

 fig. 1-lc. Bouvier 1906b:3; 1908:80. de Man 

 1911:7 



Hymenopenaeus doris. Burkenroad 1936:104; 

 1938:60. Crosnier and Forest 1973:256, fig. 83d. 



Aliporus doris. del Solar C. 1972:4. 



Material 



MEXICO— Territorio de Baja California: 1 9, USNM, 

 off Punta Chivato, Golfo de California, 1,567 m, 20 March 1889, 

 Albatross stn 3009. 1 2 , AMNH, 54 km off Punta Arena, 

 mouth of Golfo de California, 914 m, 29 April 1936, Temple- 

 ton Crocker Expedition stn 159 T-3 [station data from Beebe 

 1937]. 



COSTA RICA— 4 2 syntypes, MCZ 4648, off Cabo Velas, 

 4,082 m, 8 April 1891, Albatross stn 3414. 1 9, USNM, off 

 Cabo Velas, 4,082 m, 8 April 1891, Albatross stn 3414. 



Description. -Rostrum (Figure 17) relatively 

 short, its length about 0.2 that of carapace, reach- 

 ing between base and midlength of second anten- 

 nular article, upturned, tapering to sharp tip, and 

 with ventral margin straight. Rostral plus epi- 

 gastric teeth 7-8, sharp; epigastric tooth situated 

 about 0.4 cl from orbital margin, first rostral tooth 

 (largest of all) at approximately 0.3, and base of 



FIGURE 17. — Hymenopenaeus doris. syntype S 32.5 mm cl, off Cabo Velas, Costa Rica. Cephalothorax, lateral view. 



283 



