U.S. DEPARTMENT OF COMMERCE 



Juanita M. Kreps, Secretary 



NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



Richard A. Frank, Administrator 



NATIONAL MARINE FISHERIES SERVICE 



Robert W. Schoning, Director 



Fishery Bulletin 



The Fishery Bulletin carries original research reports and technical notes on investigations in fishery science, engineering, and 

 economics. The Bulletin of the United States Fish Commission was begun in 1881; it became the Bulletin of the Bureau of Fisheries in 

 1904 and the Fishery Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through volume 46; the last 

 document was No. 1103. Beginning with volume 47 in 1931 and continuing through volume 62 in 1963, each separate appeared as a 

 numbered bulletin. A new system began in 1963 with volume 63 in which papers are bound together in a single issue of the bulletin 

 instead of being issued individually. Beginning with volume 70, number 1, January 1972, the Fishery Bulletin became a periodical, 

 issued quarterly. In this form, it is available by subscription from the Superintendent of Documents, U.S. Government Printing Office, 

 Washington, DC 20402. It is also available free in limited numbers to libraries, research institutions, State and Federal agencies, and 

 in exchange for other scientific publications. 



EDITOR 



Dr. Bruce B. Collette 



Scientific Editor, Fishery Bulletin 



National Marine Fisheries Service 



Systematics Laboratory 



National Museum of Natural History 



Washington, DC 20560 



Editorial Committee 



Dr. Elbert H. Ahlstrom 



National Marine Fisheries Service 



Dr. William H. Bayliff 



Inter-American Tropical Tuna Commission 



Dr. Roger F. Cressey, Jr. 

 U.S. National Museum 



Mr. John E. Fitch 



California Department of Fish and Game 



Dr. William W. Fox, Jr. 

 National Marine Fisheries Service 



Dr. Marvin D. Grosslein 

 National Marine Fisheries Service 



Dr. Edward D. Houde 

 University of Miami 



Dr. Merton C. Ingham 



National Marine Fisheries Service 



Dr. Reuben Lasker 



National Marine Fisheries Service 



Dr. Sally L. Richardson 

 Oregon State University 



Dr. Paul J. Struhsaker 



National Marine Fisheries Service 



Dr. Austin Williams 



National Marine Fisheries Service 



Kiyoshi G. Fukano, Managing Editor 



The Fishery Bulletin is published quarterly by Scientific Publications Staff, National Marine Fisheries Service, NOAA, Room 450, 

 1 107 NE 45th Street, Seattle, WA 98105. Controlled circulation postage paid at Tacoma, Wash. 



The Secretary of Commerce has determined that the publication of this periodical is necessary in the transaction of the public 

 business required by law of this Department Use of funds for printing of this periodical has been approved by the Director of the 

 Office of Management and Budget through 31 December 1978. 



