U.S. Department 

 of Commerce 



Carios M. Gutierrez 



Secretary 



National Oceanic 

 and Atmospheric 

 Administration 



Vice Admiral 



Conrad C. Lautenbacher Jr., 



USN (ret.) 



Under Secretary for 

 Oceans and Atmosphere 



National Marine 

 Fisheries Service 



William T. Hogarth 



Assistant Administrator 

 for Fisheries 



Scientific Editor 



Adam Moles, Ph.D. 



Associate Editor 



Elizabeth Siddon 



Ted Stevens Marine Research Institute 

 Auke Bay Laboratories 

 Alaska Fisheries Science Center 

 17109 Pt. Lena Loop Road 

 Juneau, Alaska 99801 



^^«>*TOFCo^ 



*'>irEs o* 



The Fishery Bulletin (ISSN 0090-0656) 

 is published quarterly by the Scientific 

 Pubhcations Office, National Marine 

 Fisheries Service, NOAA, 7600 Sand 

 Point Way NE, BIN C15700. Seattle, WA 

 98115-0070. Periodicals postage is paid 

 at Seattle, WA. POSTMASTER: Send 

 address changes for subscriptions to Fish- 

 ery Bulletin, Superintendent of Docu- 

 ments, Attn.: Chief, Mail List Branch, 

 Mail Stop SSOM> Washington, DC 20402- 

 9373. 



Although the contents of this publica- 

 tion have not been copyrighted and may 

 be reprinted entirely, reference to source 

 is appreciated. 



The Secretary of Commerce has deter- 

 mined that the publication of this peri- 

 odical is necessary according to law for 

 the transaction of public business of this 

 Department. Use of funds for printing of 

 this periodical has been approved by the 

 Director of the Office of Management and 

 Budget. 



For sale by the Superintendent of 

 Documents, U.S. Government Printing 

 Office, Washington, DC 20402. Subscrip- 

 tion price per year: $36.00 domestic and 

 $50.40 foreign. Cost per single issue: 

 .$28.00 domestic and $35.00 foreign. See 

 back for order form. 



Managing Editor 



Sharyn Matriotti 



National Marine Fisheries Service 

 Scientific Publications Office 

 7600 Sand Point Way NE, BIN C15700 

 Seattle, Washington 98115-0070 



Editorial Committee 



Jeffrey M. Leis Australian Museum, Sydney, Australia 



Thomas Shirley Texas A&M University 



David Somerton National Manne Fisheries Service 

 Mark Terceiro National Marine Fisheries Service 



Fishery Bulletin web site: www.fishbull.noaa.gov 



The Fishen Bulletin carries original research reports and technical notes on investigations in 

 fishery science, engineering, and economics. It began as the Bulletin of the United States Fish 

 Commission in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery 

 Bulletin of the Fish and Wildhfe Service in 1941. Separates were issued as documents through 

 volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and continuing 

 through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system 

 began in 1963 with volume 63 in which papers are bound together in a single issue of the 

 bulletin. Beginning with volume 70, number 1, January 1972, the Fishery Bulletin became a 

 periodical, issued quarterly. In this form, it is available by subscription from the Superintendent 

 of Documents, U.S. Government Printing Office, Washington, DC 20402. It is also available free in 

 limited numbers to libraries, research institutions, State.and Federal agencies, and in exchange 

 for other scientific publications. 



