810 



Fishery Bulletin 97(4), 1999 



100 



o 



study of this species (Gabr, et 

 al., 1998) is quite likely, 

 partly because the spawning 

 season is greatly protracted 

 (Fig. 3). Indeed, it appears 

 that this species is capable of 

 meeting the demands of oo- 

 cyte production without me- 

 tabolizing reserves of other 

 tissues. Both males and fe- 

 males continue to feed as ma- 

 turity is reached, thus energy 

 for gonad production is di- 

 verted primarily from the 

 food supply, although some 

 diversion from somatic tissue 

 may also occur. In the context 

 of this paper and results from 

 S. doll fust (Gabr et al., in 

 press), variation in reproduc- 

 tive strategies correlates with variations in feeding 

 and growth patterns as suggested previously for 

 other coleoids (Mangold et al., 1993). 



Acknowledgments 



We are especially grateful to Emma Hatfield for her 

 help and advice about data interpretation during 

 preparation of the manuscript. We also wish to ex- 

 press our thanks to Michael Maxwell, Paul Rago, 

 Jean Boal, and Nadav Shashar for frequent advice 

 and assistance in statistical analysis. This study, part 

 of a doctoral dissertation, was supported by the Em- 

 bassy of the Arab Republic of Egypt, Cultural and 

 Educational Bureau. A portion of this study was sup- 

 ported by the British Council through Liverpool 

 University and the Marine Biology Department, Suez 

 Canal University. 



Female 



Male 



100 



o 



I u in 



Maturity stages 



I II III IV 



Maturity stages 



Figure S 



Stomach fullness for each maturity stage of both sexes of Sepia pharaonis. = empty; 

 1 = one-quarter full: 2 = half full; 3 = three-quarters full; 4 = full or distended). 



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