1968; McNeil, Lawton, 1970; Klekowski, 1970; Hughes, 1970; Bregman, 

 1971; and others). After calculating the value of K2, it is not 

 difficult to determine the production of individuals of various sizes 

 (weights), then the production of the population, as the sum of the 

 growth of all individuals present in the population during the time of 

 the analysis. Obviously, to analyze the annual production of a 

 population by the ecophysiologic method, it is also necessary to know 

 the relationship of the numbers of individuals in cohorts during various 

 seasons of the year, particularly since the rate of elimination and 

 intensity of metabolism and growth of individuals change greatly from 

 season to season. 



As we can see, all of the methods analyzed above for calculation of 

 production require regular, year-round observations of changes in the 

 numbers of individuals of various ages. These observations of 

 populations of benthic organisms are frequently impossible, particularly 

 in areas far from scientific research institutions. Considering this 

 fact, as well as the difficulty of tracing the dynamics of the 

 populations of organisms in the open seas, a static-dynamic method of 

 determination of production has been suggested, based on analysis of the 

 size-weight and age structure of populations at the moment of 

 observation as a result of the annual production process (Golikov, 

 1970). The method is based on the fact that the structure of a 

 population is used to establish the growth rate and age of individuals 

 (with a check on the basis of morphologic indications of growth, if 

 there are such), after which the quantity of living matter produced by 

 the population during the year before the time of analysis of its 

 initial state (growth production of population Pg) is calculated by 

 estimating the weight gain of all individuals detected at the moment of 

 observation, using the equation: 



Pg = I N,(W,+i - W,) ^ (4.4) 



T=0 



or the classical equation (4.2). 



The value of At in the version analyzed is equal to one year. 

 Based on equation (4.4), the weight gain of all individuals surviving 

 and those eliminated during the time period analyzed is calculated, 

 while equation (4.2) is used to determine the weight gain of all 

 individuals surviving and half of those eliminated during this period of 

 time. Obviously, with a sharp decrease in the population of the 

 generations at the beginning of the time sector analyzed, with 

 relatively uniform elimination, and in those cases when the time of 

 elimination of individuals is not known, the best result will be yielded 

 by an estimate of the growth production using the equation generally 

 used for this purpose (4.2). However, in those cases when most of the 

 elimination occurs at the end of the time sector analyzed (for example, 

 elimination after spawning), calculation of growth production by 

 equation (4.4) may be more accurate. 



Determination of growth production of a population using a single 

 sample per year is based, as in calculation of production by the other 



309 



