U.S. DEPARTMENT OF COMMERCE 



Robert Mosbacher, Secretary 



National Oceanic and Atmospheric Administration 



John A. Knauss, Under Secretary for Oceans and Atmosphere 



National Marine Fisheries Service 



William W. Fox, Jr., Assistant Administrator for Fisheries 



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B U 



T I N 



The Fishery Bulletin carries original research 

 reports and technical notes on investigations in 

 fishery science, engineering, and economics. 

 The Bulletin of the United States Fish Commis- 

 sion was begun in 1881; it became the Bulletin 

 of the Bureau of Fisheries in 1904 and the 

 Fishery Bulletin of the Fish and Wildlife Ser- 



vice in 1941. Separates were issued as docu- 

 ments through volume 46; the last document 

 was No. 1103. Beginning with volume 47 in 1931 

 and continuing through volume 62 in 1963, 

 each separate appeared as a numbered bulle- 

 tin. A new system began in 1963 with volume 63 

 in which papers are bound together in a single 



issue of the bulletin instead of being issued in- 

 dividually Beginning with volume 70, number 

 1, January 1972, the Fishery Bulletin became 

 a periodical, issued quarterly It is available 

 free in limited numbers to libraries, research 

 institutions. State and Federal agencies, and 

 in exchange for other scientific publications. 



Scientific Editor 



Dr. Andrew E. Dizon 



Southwest Fisheries Center 



Lajolla Laboratory 



National Marine Fisheries Service, NOAA 



P.O. Box 271, Lajolla, CA 92038 



Editorial Committee 



Dr. Jay Barlow 



National Marine Fisheries Service 



Dr. William H. Bayiiff 



Inter-American Tropical Tuna Commission 



Dr. George W. Boehlert 

 National Marine Fisheries Service 



Dr. Bruce B. CoUette 



National Marine Fisheries Service 



Dr. Robert C. Francis 

 University ofWashington 



Dr. James R. Kitchell 

 University of Wisconsin 



Dr. William J. Richards 

 National Marine Fisheries Service 



Dr. Tim D. Smith 



National Marine Fisheries Service 



Managing Editor 



Mary S. Fukuyama 



The Fishery Bulleiin ( ISSN 0090-0656) is published quarterly by the Scientific Publications Office, National Marine Fisheries Service, NOAA, 7600 Sand Point Way NE, BIN 



C15''00, Seattle, WA981 15, Second class postage.is paid In Seattle. WA, and additional offices. POSTMASTER send address changes for subscriptions to Fisfiery Bulletin, 



Superintendent of Documents, U.S. Government Printing Office, Washington, DC 20402. 



Although ;the contents have not been copyrighted and may be reprinted entirely reference to source is appreciated. 



The Secretary of Commerce has determined that the publication of this periodical is necessary in the transaaion of the public business required by law of this 



Department, Use of funds for printing of this periodical has been approved by the Director of the Office of Management and Budget. 



For sale by the Superintendent of Documents, US, Government Printing Office, Washington, DC 20402-Subscription price per year: $16.00 domestic and $20.00 foreign. 



Cost per single issue: $9.00 domestic and $11 25 foreign. 



