FISHERY BULLETIN: VOL. 87, NO. 1 



we paraphrase essential features of A. Milne Ed- 

 wards's original description. 



Near Calappa and Mursia; distinguished from 

 former because carapace not extended above ambu- 

 latory legs (Fig. 7) in manner of a shield, and from 

 latter by absence of large spines laterally prolonged 

 beyond cephalothoracic shield; front very narrow 

 and ornamented with 2 small slightly divergent 

 points very similar to those of Calappa; [orbital] 

 border cut by two narrow fissures. 



Figure l.—Calappilia brooksi Ross and Scolaro, carapace and left 

 eyestalk; USNM 648599, Upper Eocene, Fla. 



Carapace very convex, recalling that of Calappa 

 or certain representative Leucosiidae; gastric and 

 cardiac regions separated in lateral portions by deep 

 grooves; hepatic region not clearly delimited; bran- 

 chial region very inflated in anterior part but much 

 narrowed posteriorly, surface covered with coarse 

 tubercles in anterior part; posterior branchial lobe 

 extended, constituting a prominence directed 

 laterally and a little posteriorly; posterior border 

 bearing a tubercle much less developed than bran- 

 chial prominence at level of branchiocardiac groove. 



Ambulatory legs missing; fragment of chela with 

 very compressed dactyl bearing granular crest, 

 armed at base with large tubercle recalling that 

 developed in Calappa; palm covered with large 

 tubercles analogous to those ornamenting cara- 

 pace, and their size notable compared to those on 

 body. 



Measurements of selected species in mm. — Cara- 

 120 



pace: C. brooksi length 18.8, width 21.5; C. dacica 

 length 32, width 37; C. hondoensis length 19, width 

 18.7. 



Remarks. — The features of Calappilia mentioned 

 by A. Milne Edwards suggest much closer similar- 

 ity to Calappa than to Paracyclois, and the brief 

 diagnosis by Rathbun (1930) confirms this in broad 

 outline. All of the species of Calappilia are small, 

 comparing favorably with the range of sizes shown 

 by the two species of Cyclozodion described here. 

 There is considerable diversity in ornamentation of 

 the carapace among species of Calappilia, with a 

 tendency to development of coarse tubercules dor- 

 sally and along the margins, especially postero- 

 laterally, but minimal development of posterolateral 

 winglike projections, with some exceptions. Lobular 

 tubercles along this margin are usually similar in 

 size, although in C. scopuli (Quayle and Collins 

 1981:740, pi. 104, fig. 8) there is a developed pos- 

 terolateral spine and, except for the problematic 

 frontoorbital region, a marked similarity to Cyclo- 

 zodion in outline of the carapace. The holotype of 

 Calappilia hondoensis (USNM 371094) has an ob- 

 scure posterolateral spine rather wider than long. 

 Rathbun (1930) pointed out that Milne Edwards's 

 (1873) figure of C. verrucosa is longer than wide 

 whereas the measurements given show it wider than 

 long. The left eyestalk of C. brooksi (USNM 648599, 

 Fig. 7), fossilized projecting forward in its orbit, 

 seems relatively slender compared with eyestalks 

 of both Cyclozodion and Paracyclois, although only 

 a remnant of it may be preserved. 



On the basis of size, shape, and ornamentation of 

 the carapace, relative thickness of eyestalks, and 

 age, we regard Early Tertiary Calappilia and 

 Recent Paracyclois as distinct. Calappilia scopuli 

 and perhaps C. hondoensis seem to form closer 

 links with Recent Cyclozodion than with Calappa, 

 emphasizing similarities among the latter three 

 genera. 



ACKNOWLEDGMENTS 



We thank A. B. Johnson of MCZ for loan of 

 material, R. E. Gibbons for drafting the graph, and 

 G. A. Bishop, B. B. CoUette, R. B. Manning, and 

 N.N. Rabalais for critical review of the manuscript. 



LITERATURE CITED 



Chace, F. a., Jr. 



1939. Reports on the scientific results of the first Atlantis 

 Expedition to the West Indies, under the joint auspices of 



