LAROCHE and RICHARDSON: REPRODUCTION OF NORTHERN ANCHOVY 



fall and winter, during the peak spawning period, 

 and return inshore in spring. Huppert et al. (foot- 

 note 3) indicated similar movements but provided 

 little additional information. Brewer 11978) 

 suggested that mature northern anchovies in the 

 vicinity of San Pedro Bay move to deeper, cooler 

 waters offshore to spawn. The distinct geographic 

 segregation of mature and immature northern an- 

 chovies in the northern subpopulation which oc- 

 curs because of an offshore migration of spawning 

 fish may represent an additional racial difference 

 among the three northern anchovy subpopula- 

 tions in the northeast Pacific. 



ACKNOWLEDGMENTS 



Wayne A. Laroche took the photomicrographs of 

 oocytes. Jerome D. Spratt and John S. Sunada of 

 the California Department of Fish and Game 

 checked our otolith readings and provided useful 

 advice concerning interpretation of northern an- 

 chovy age data. Percy L. Donaghay gave us many 

 helpful ideas for data analyses and Rae Deane 

 Leatham generously provided assistance with all 

 computer work. David L. Stein rendered a careful 

 review of the manuscript. John R. Hunter, Na- 

 tional Marine Fisheries Service (NMFS), La Jolla, 

 read an earlier version of the manuscript and 

 suggested useful ways to improve it. The following 

 individuals helped us during this study by provid- 

 ing information on the location of northern an- 

 chovies at sea or collected and saved fish for us: P. 

 Killian, W. Nichols, and G. Muhlberg of the Clat- 

 sop Community College, Astoria, Oreg.; M. Hosie, 

 G. Hettman, J. Lukas, and B. Forsberg of the Ore- 

 gon Department of Fish and Wildlife; T. Durkin, 

 NMFS, Hammond, Oreg.; W Pearcy, D. Stein, and 

 E. Krygier of the Oregon State University, Corval- 

 lis, Oreg. 



This work is a result of research sponsored by 

 the Oregon State University Sea Grant College 

 Program, supported by NOAA Office of Sea Grant, 

 Department of Commerce, under Grant No. 04-6- 

 158-44094. 



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