U.S. DEPARTMENT OF COMMERCE 



Malcolm Baldrige, Secretary 



NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



Terry L. Leitzell, Assistant Administrator for Fisheries 



NATIONAL MARINE FISHERIES SERVICE 



Fishery Bulletin 



The Fishery Bulletin carries original research reports and technical notes on investigations in fishery science, engineering, and 

 economics. The Bulletin of the United States Fish Commission was begun in 1881; it became the Bulletin of the Bureau of Fisheries in 

 1904 and the Fishery Bulletin of the Fish and Wildlife Service in 1941 . Separates were issued as documents through volume 46; the last 

 document was No. 1103. Beginning with volume 47 in 1931 and continuing through volume 62 in 1963, each separate appeared as a 

 numbered bulletin. A new system began in 1963 with volume 63 in which papers are bound together in a single issue of the bulletin 

 instead of being issued individually. Beginning with volume 70, number 1, January 1972, theFishery Bulletin became a periodical, 

 issued quarterly. In this form, it is available by subscription from the Superintendent of Documents, U.S. Government Printing Office, 

 Washington, DC 20402. It is also available free in limited numbers to libraries, research institutions. State and Federal agencies, and 

 in exchange for other scientific publications. 



EDITOR 



Dr. Jay C. Quasi 



Scientific Editor, Fishery Bulletin 



Northwest and Alaska Fisheries Center 



Auke Bay Laboratory 



National Marine Fisheries Service, NOAA 



P.O. Box 155, Auke Bay, AK 99821 



Editorial Committee 



Dr. Bruce B. Collette Dr. Reuben Lasker 



National Marine Fisheries Service National Marine Fisheries Service 



Dr. Edward D. Houde Dr. Jerome J. Pella 



Chesapeake Biological Laboratory National Marine Fisheries Service 



Dr. Merton C. Ingham Dr. Sally L. Richardson 



National Marine Fisheries Service Gulf Coast Research Laboratory 



Kiyoshi G. Fukano, Managing Editor 



The Fishery Bulletin (USPS 090-870) is published quarterly by Scientific Publications Office, National Marine Fisheries Service, NOAA, 

 Room 336, 1 700 Westlake Avenue North, Seattle, WA 98109. Controlled circulation paid to Finance Department, USPS, Washington, 

 DC 20260. 



Although the contents have not been copyrighted and may be reprinted entirely, reference to source is appreciated. 



The Secretary of Commerce has determined that the publication of this periodical is necessary in the transaction of the public business 

 required by lav*? of this Department. Use of funds for printing of this periodical has been approved by the Director of the Office of 

 Management and Budget through 31 March 1982. 



