FISHERY BULLETIN: VOL. 78, NO. 3 



Table 8. — Summary of additional studies in which juvenile and adult Engraulis mordax from the northern subpopulation were 



collected. 



General area 



Dates 



Principal gear 



Sampling effort 



Reference 



Washington coast between Cape 8-18 Nov 1 966 



Flattery and Destruction 



Island (20-100fm) and 



between Grays Harbor and 



Columbia River (10-50 fm) 

 Washington-Oregon coast 



between Cape Flattery and 



Yaquina Bay 



18 Nov- 16 Dec. 1966 and 

 3 Jan -8 Apr. 1967 



Washington-Oregon coast 

 between Cape Flattery and 

 HecetaHead (15-100 fm) 



Tillamook Bay, Oreg. 



Yaquina Bay, Oreg. 



Coos Bay, Oreg. 



Humboldt Bay, Calif. 



1 5 IVIay-2 June 1967 



May 1974-May 1976 

 July 1964-Sept. 1967 

 June-Sept. 1970 



Apr. 1974-Oct. 1976 



fish schools) 

 2/3 scale Cobb pelagic trawl 

 (echosounder used to locate 

 fish schools) 



Standard Cobb pelagic trawl, 

 2/3 scale Cobb pelagic 

 trawl, and 2 experimental 

 anchovy trawls (echosounder 

 used to locate fish schools) 

 BCF Universal trawl (echo- 

 sounder used to locate fish 

 schools) 

 6 m try net; 46 m beach seine 

 6 m otter trawl (plus other gear) 

 61 m beach seine; 30 m bag 



seine (plus other gear) 

 Echosounder — to determine 

 distribution of anchovy 

 schools; 200 m lampara bait 

 seine 66.7 x 6.7 m purse 

 seine (plus other gear) 



2 tows 



71 tows (most tows made 

 in Jan-Apr.) 



7 tows 



-biweekly, then monthly 



^monthly 



Not specified 



Weekly echosounder 

 surveys plus 45 net 

 hauls 



BCF exploratory cruise no. 82, 

 RV John N. Cobb^ 



BCF gear research cruise 

 no. 8, MV Baron^ 



BCF exploratory cruise no. 87, 

 RV John N. Cobb^ 



Forsberg et a!.-' 

 Beardsley (1969) 

 Cummings and Schwartz^ 



Waldvogel(1977) 



'Barraclough, W. E, D. G. Robinson, and J. D. Fulton. 1968. Data record — Number, size composition, weight, and food of larval and juvenile fish caught with a 

 two-boat surface trawl in Saanich Inlet April 23-July 21, 1968. Fish. Res. Board Can., Manuscr. Rep. Ser. 1004, 305 p. 



2Robinson, D. G. 1969. Data record — Number, size composition, weight and food of larval and juvenile fish caught with a two-boat surface trawl in the Strait of 

 Georgia July 4-6, 1967. Fish. Res. Board Can., Manuscr, Rep Ser. 1012, 71 p. 



^Information from cruise reports. Northwest and Alaska Fisheries Center, NMFS, NOAA, 2725 Montlake Blvd. East, Seattle, WA 98112. 



■•Forsberg, B. O, J. A, Johnson, and S. M. Klug. 1977. Identification, distribution, and notes on food habits of fish and shellfish in Tillamook Bay Oregon. Oreg. 

 Dep. Fish Wildl. Res. Sec, Fed. Aid Prog, Rep. Fish. 1977, 117 p. 



^Cummings, E., and E. Schwartz. 1971. Fish in Coos Bay, Oregon, with comments on distribution, temperature, and salinity of the estuary Oreg. Dep. Fish 

 Wildl., Coastal rivers invest— -Inf. Rep. 70-11, 22 p. 



in spawning condition were found offshore with 

 main concentrations between lat. 43° and 47° N 

 and -65-157 km offshore. They occurred in small 

 schools near the surface at night and deeper in the 

 water column during daylight. Immature fish 

 (mostly <100 mm SL) remained in nearshore 

 coastal areas or in bays. They were taken in Grays 

 Harbor, Wash., around the Columbia River mouth, 

 in Tillamook Bay, Yaquina Bay, and Coos Bay, 

 where they were observed feeding at the surface 

 during daylight. In Humboldt Bay, mature fish left 

 in June and July leaving only immature fish in the 

 bay through the summer. Adults presumably 

 moved offshore to spawn, although it is not known 

 whether they moved north off Oregon and Wash- 

 ington or elsewhere. Adults returned to the bay 

 around mid-September in spent condition. 



In fall, October through December, northern an- 

 chovies are no longer abundant around British 

 Columbia. The fishery of the 1940's was generally 

 not in operation that season. Off Washington and 



Oregon, adults eventually return to coastal waters 

 from offshore spawning areas although few adults 

 were collected in the fall. Fall catches occurred 

 mainly off the Columbia River mouth (13-61 m 

 depth) and Grays Harbor (35-37 m depth). Imma- 

 ture fish appeared to leave the bays and estuaries 

 and returned to nearshore coastal waters. Young 

 fish were seen in the Columbia River in October, 

 feeding at the surface. No anchovies were collected 

 in Tillamook Bay, Yaquina Bay, or Coos Bay. 

 Juveniles and adults left Humboldt Bay in late 

 October- November. 



A pronounced and well-defined onshore-offshore 

 segregation of mature and immature fish and in- 

 ferred offshore spawning migration during the 

 spawning period as observed in the northern sub- 

 population off Washington, Oregon, and northern 

 California has not been documented for northern 

 anchovies in the central or southern subpopula- 

 tions. Baxter (1967) stated that off California 

 northern anchovies apparently move offshore in 



616 



