COLLETTE and RUSSO: SPANISH MACKERELS 



1853:42 (India). Giinther 1860:370 (descrip- 

 tion; Malaya). Bleeker 1861a:52 (Singapore; 

 listed). Bleeker 1861b:74 (Pinang, Malaya; 

 listed). Day 1873:225 (description, range). 

 Day 1878:256 (description). 



Scomberomorus lineolatum. Malpas 1926:74 (3 

 males, 54.5-82.5 cm TL, 1.1-3.4 kg, Ceylon). 

 Frost 1928:329 (otolith similar to that of S. 

 regalis ). 



Scomberomorus lineolatus. Munro 1943:68, 70 

 (placed in subgenus Indocybium ; vertebral 

 count of 21 + 29 = 50 pertains to another species 

 of Scomberomorus, such as S. guttatus). De 

 Beaufort 1951:235-236 (synonymy, description, 

 range). Tham 1953:49 (Singapore Straits), 50 

 (correlation of catch with physical factors and 

 presence of food fishes such as Stolephorus and 

 Clupea). Scott 1959:114 (description; Malaya), 

 photograph. Bauchot and Blanc 1961:372 

 (types of Cybium lineolatum ). Jones 1962:117- 

 119 (eggs, larvae, and juveniles). Jones and 

 Silas 1961:197-198 (description, range), fig. 4 

 (680 mm adult), fig. 5C (not 5D as legend reads, 

 lateral view of head), fig. 6D (gill arch), fig. 7E 

 (caudal peduncle keels). Jones and Silas 1964: 

 58-61 (synonymy, description, range), pi. 7, fig. 

 A (photograph of 740 mm specimen). Silas 

 1964:317, 323-324 (specimens from India only). 

 Jones and Kumaran 1964:347 (larvae as yet 

 undescribed). Blanc and Bauchot 1964:447 

 (types of Cybium lineolatum ), pi. Ill, figs. 15, 16 

 (photographs of type-specimens). Rao 1964: 

 592-594 (teleosts constitute 97% of diet of ju- 

 veniles, Waltair coast of India). Thiemmedh 

 1966:140 (common names). Collette 1966:367- 

 368 (type of Cybium lineolatum). Tongyai 

 1966a:7-10 (synonymy, occurrence, Thailand), 

 pi. 2D. Tongyai 1966b:3-15 (5 specimens, 46.5- 

 76.5 cm FL; Terutao Is., Andaman Sea). Pa- 

 thansali 1968:1002-1003 (fishery; Malaya). Si- 

 las 1968:1114 (fishery; Gulf of Mannar), pi. 3A 

 (photograph of adult). Rajan et al. 1969:90 

 (outer channel; Chilka Lake, India). Tongyai 

 1970:559-561 (found in areas of low turbidity 

 and high salinity offshore; Thailand). Merce- 

 ron 1970:72 (specimens from near Sihanouk- 

 ville, Cambodia). Tongyai 1971a:16-18 (Thai- 

 land). Fernando 1972:524, 530 (incidental 

 catches in trawls; Wadge Bank, Ceylon). Rich- 

 ards and Klawe 1972:14 (range), 92 (references 

 to larvae and juveniles). Banerji 1973:129-130 

 (seerfish fishery; India). Magnuson 1973:350 

 (short pectoral fin). Orsi 1974:175 (listed; Viet- 

 nam). Tham 1974 (possible predator of Stole- 



phorus). Shiino 1976:231 (common name). 

 *Rao and Ganapati 1977:101-111 (postlarvae 

 and juveniles; India). Klawe 1977:2 (common 

 name, range). *Devaraj 1977 (osteology). 

 Collette 1979:29 (characters). Collette and 

 Russo 1979:13 (diagnostic characters, range). 

 *Devaraj 1982 (age and growth). Cressey et 

 al. 1983:264 (host-parasite list, 3 copepod spe- 

 cies). Collette and Nauen 1983:68 (description, 

 range), fig. 



Scomberomorus guttatus. Not of Bloch and 

 Schneider 1801. Fraser-Brunner 1950:160 (Cyb- 

 ium lineolatum placed in synonymy of Scomber- 

 omorus guttatus ). 



Indocybium lineolatum. Munro 1955:221 (de- 

 scription; Ceylon); fig. 651. Chacko et al. 1968: 

 1006 (fishery; Madras). 



Types. — Holotype: MNHN A.6866; Malabar coast 

 of India; Dussumier; 707 mm FL; D about XVII + 

 17 + IX; A 19 + X; Pj 21; RGRj 2+1 + 8=11; pattern 

 of three rows of elongate streaks still visible on 

 type in 1975. Photograph of type published by 

 Blanc and Bauchot (1964:pl. 3, fig. 15). Paratype: 

 MNHN 6357; Mahe (Malabar Coast), Belenger; 

 only head and tail of a fish about 710 mm FL 

 (judging from head length of 145 mm). Photograph 

 of paratype published by Blanc and Bauchot 

 (1964:pl. 3, fig. 16). 



Diagnosis. — The only species of Scomberomorus 

 that has a pattern of short lines on its sides (Fig. 

 57). Other species have some spots, blotches, or 

 bars, or are plain. Posterior end of maxilla greatly 

 expanded as in S. plurilineatus and S. semifascia- 

 tus. Anterior end of premaxilla forms an acute 

 angle (Fig. 22b). Ascending process of premaxilla 

 very long as in S. sinensis and Acanthocybium. 

 Supracleithrum wide (Fig. 41a), 53-57% of length, 

 as in S. niphonius. Foramen between last pectoral 

 radial and coracoid larger than in any other 

 species of Scomberomorus. 



Description. — Lateral line gradually descending 

 to midline on caudal peduncle. Intestine with two 

 folds and three limbs (Fig. 3g). Spines in first 

 dorsal fin 15-18, usually 16 or 17 (Table 9); second 

 dorsal fin rays 15-22, usually 17 or 18 (Table 10); 

 dorsal finlets 7-10, usually 9 (Table 10); anal fin 

 rays 17-22, usually 20 (Table 11); anal finlets 7-10, 

 usually 9 or 10 (Table 11); pectoral fin rays 20-24, 

 usually 23 (Table 12). Precaudal vertebrae 18-20, 

 usually 19 (Table 6); caudal vertebrae 25-28, 

 usually 27 (Table 7); total vertebrae 44-46, usually 



639 



