COLLETTE and RUSSO: SPANISH MACKERELS 



gonads; mature in June), fig. 7 (oocytes). Pos- 

 tel 1958:107-111 (summary of Postel 1955a). 

 Postel 1959:163 (listed). Postel 1960:257 (Cap 

 Blanc to Senegambie). Marchal 1961:106 (lat. 

 9°12'N, long. 13°42'W). Franca 1964:table 3 

 (Angola). Daget and litis 1965:280-281 (Ebrie 

 and Abi lagoons, Ivory Coast), fig. 178. San- 

 ches 1966:146 (Angola). Blache et al. 1970:375 

 (in key), fig. 960 (not fig. 961). Conand 1970:40 

 (distribution of larvae). 



Apolectus immunis Bennett 1831:146 (original 

 description; "Atlantic Coast of North Africa"). 



Cybiummaculatum. Not of Mitchill 1815. Stas- 

 sano 1890:44 (Spanish Sahara). Vinciguerra 

 1890:100-103 (synonymy). Osorio 1898:197 

 (Sao Thome). 



Scomberomorus argyreus Fowler 1905:764-765 

 (original description, "West Africa"), pi. 51. 



Scomberomorus maculatus. Not of Mitchill 1815. 

 Fowler 1936:628-629. Cadenat 1937:482 (Da- 

 kar). Scaccini 1941:19 (synonymy in part; Mau- 

 ritania). Navarro 1943:131 (Cabo Barbas and 

 Blanco, Banco Arguin, Mauritania). Tortonese 

 1949:65 (accidental in Mediterranean Sea). 

 Sanz Echeverria 1950:1-2 (sagitta compared 

 with other scombrids), pi. 1, figs. 1-4 (photo- 

 graphs of sagittae). Mather and Day 1954:182, 

 185 (Sierra Leone, Dakar, Canary Is.; S. tritor 

 not specifically distinct from S. maculatus). 

 Tortonese 1956:7 (accidental in Mediterranean 

 Sea). Poll 1959:104-106 (description; S to Baie 

 des Tigres, Angola), fig. 34. Maurin et al. 

 1970:19 (Nouakchott, NW Africa). Lozano 

 Cabo 1970:158 (Sahara coast). Fagade and 

 Olaniyan 1973:212, 220, 224 (piscivorous, feed- 

 ing largely on Ethmalosa fimbriata in Lagos). 

 Fagade and Olaniyan 1974:249 (caught in Lagos 

 Lagoon when water was brackish). Tortonese 

 1975:354-355 (description, Italy), fig. 155. 



Scomberomorus tritor. Munro 1943:67-71 (placed 

 in subgenus Scomberomorus). Irvine 1947: 

 186-187 (Accra, Ghana), fig. 108. Fraser- 

 Brunner 1950:158 (synonymy in part), fig. 27. 

 Chaine 1957:504-509 (otoliths), pi. IV (otoliths). 

 Gras 1961:583 (Lagune de Cotonou and Lac 

 Nokoue, Dahomey). Bauchot and Blanc 1961: 

 372-373 (types). Blanc and Bauchot 1964:448 

 (types), pi. IV, figs. 19-20 (photographs of types). 

 Gorbunova 1965a:54 (spawning season). Col- 

 lette 1966:367 (types). Williams 1968:436, ta- 

 ble 593 (taken from Gambia to the Congo during 

 the Guinean Trawling Survey). Collette 1970: 

 4-5 (in key; Mediterranean Sea). Richards and 

 Klawe 1972:15 (range), 96 (references to larvae). 



Miyake and Hayasi 1972:111-3 (in key), IV-10 

 (common names). Magnuson 1973:350 (short 

 pectoral fin). Klawe 1977:2 (common name; 

 range). Penrith 1978:187 (Baia dos Tigres, 

 Angola). Collette et al. 1978:274-275 (compar- 

 ison with W Atlantic species). Collette 1979: 

 29 (characters, range). Collette and Russo 

 1979:13 (diagnostic characters, range). Col- 

 lette 1981:Scombm 7 (description, range), fig. 

 Seret and Opic 1981:332-333 (description), fig. 

 Cressey et al. 1983:264 (host-parasite list, 4 

 copepod species). Collette and Nauen 1983:79 

 (description, range), fig. 



Types of nominal species. — Holotype: MNHN 

 A. 6871; Goree, Senegal; Rang; 658 mm FL; D 

 X V + 17 + VIII; A 17 + VIII; P! 21; RGRj 2 + 1 + 10 

 = 13. Paratype: MNHN A.6868; Goree; Senegal; 

 Rang; 505 mm FL. Photographs of the holotype 

 and paratype were published by Blanc and Bau- 

 chot (1964:pl. 4, figs. 19, 20). 



Scomberomorus argyreus Fowler 1905. Holo- 

 type: ANSP 11400; west coast of Africa; Dr. Sav- 

 age; 148 mm FL; D XVII + 17 + VIII; A 19 + VIII; P, 

 22-22; RGRi 2 + 1 + 11 = 14; vertebrae 18 + 28 = 46. 



Diagnosis. — This species possesses nasal denti- 

 cles as do the other five species of the regalis group 

 (brasiliensis, concolor, maculatus, regalis, and 

 sierra) but lacks the artery branching from the 

 fourth left epibranchial artery that is present in 

 the other species (Fig. 7b). Intercalar spine absent 

 as in the other five species of the regalis group and 

 S. niphonius. 



Description. — Lateral line gradually descending 

 to midline on caudal peduncle. Intestine with two 

 folds and three limbs (Fig. 3r). Spines in first 

 dorsal fin 15-18, usually 17 or 18 (Table 9); second 

 dorsal fin rays 16-19, usually 17 (Table 10); dorsal 

 finlets 7-9, usually 8 (Table 10); anal fin rays 

 17-20, usually 18 or 19 (Table 11); anal finlets 

 7-9, usually 8 (Table 11); pectoral fin rays 20-22, 

 usually 21 (Table 12). Precaudal vertebrae 18 or 19, 

 usually 19 (Table 6); caudal vertebrae 27 or 28, 

 usually 27 (Table 7); total vertebrae 46 or 47, 

 usually 46 (Table 8). Gill rakers on first arch 

 (1-3) + (10-13) =12-15, usually 2 + (11-12)= 13-14 

 (Table 5). Postel (1955a) reported a range of 10-15 

 gill rakers for 240 males and 520 females, 94% of 

 both sexes 12-14. Morphometric characters given 

 in Table 30. 



Size. — Maximum size of males 83.9 cm FL, fe- 



669 



