U.S. DEPARTMENT OF COMMERCE 



Malcolm Baldnge, Secretary 



NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



John V. Byrne, Administrator 



NATIONAL MARINE FISHERIES SERVICE 



William G. Gordon. Assistant Administrator 



Fishery Bulletin 



The Fishery Bulletin carries original research reports and technical notes on investigations in fishery science, engineering, and 

 economics. The Bulletin of the United Stales Fish Commission was begun in 188), it became the Bulletin of the Bureau of Fisheries 

 in 19dl and the Fishery Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through volume 46; the 

 last document was No. 1 103. Beginning with volume 47 in 1931 and continuing through volume 62 in 1963, each separate appeared as 

 a numbered bulletin. A new system began in 1963 with volume 63 in which papers are bound together in ,. single issue of the bulletin 

 instead of being issued individually. Beginning with volume 70. number 1. January 1972. the Fishery Bulletin became a periodical, 

 issued quarterly. In this form, it is available by subscription from the Superintendent of Documents. U.S. Government Printing 

 Office. Washington DC 20402. It is also available free in limited numbers to libraries, research institutions. State and Federal 

 agencies, and in exchari nl ific publications 



SCIENTIFIC EDITORS. Fishery Bulletin 



Dr. Carl J. Sindermann 



Northeast Fisheries Center Sandy Hook Laboratory 



National Marine Fisheries Service, NOAA 



Highlands, NJ 07732 



Dr. William J. Richards 



Southeast Fisheries Center Miami Laboratory 



National Marine Fisheries Service, NOAA 



Miami, FL 33149-1099 



Editorial Committee 



Dr. Bruce B. Collette 



National Marine Fisheries Service 



Dr. Edward D. Houde 

 Chesapeake Biological Laboratory 



Dr. Merton C. Ingham 



National Marine Fisheries Service 



Dr. Reuben Lasker 



National Marine Fisheries Service 



Dr. Donald C. Malins 



National Marine Fisheries Service 



Dr. Jerome J. Pella 



National Marine Fisheries Service 



Dr. Jay C. Quast 



National Marine Fisheries Service 



Dr. Sally L. Richardson 



Gulf Coast Research Laboratory 



Mary S. Fukuyama, Managing Editor 



■\ Bulletin - published quarterly by the Scientific Publications Office, National Marine 



iinl Way NE, BIN C15700, Seattle, WA 98115. Second class postage is paid at Seattle. Wash., and 

 additional offices. POSTMASTER send addre - for subscriptions to Superintendent of Docui S Government 



Printing Office. Washington, DC 20 10 



Although the contents have nm been copyrighted and may be reprinted entirely, reference to source is appreciated. 



The - mmerce has determined that the publication of this periodical is necessary in the transaction of the public 



iiss required by law of this Department. I is for printing of this periodical has been approved by the Director o( the 



Office of Management and Budget through 1 April 19 



