Brouwer and Griffiths: Reproductive biology of Argyrozona argyrozona 



263 



100 

 90 

 80 

 70 

 60 

 50 

 40 

 30 

 20 

 10 

 



100 

 90 

 80 

 70 

 60 

 50 

 40 

 30 

 20 

 10 

 



Male 



m m m 



Saj 



1 



A BS 



12 3 4 5 



9 10 



11 12 



12 3 4 5 



6 7 



Months 



10 11 12 



n=998 



□ 1 



□ 3 



□ 4 



Figure 4 



Monthly variation in the proportion of macroscopic gonad stages of carpenter 

 {Argyrozona argyrozona) >L 50 caught in the Tsitsikamma National Park (March 1996-July 

 1999). Numbers in the legend refer to the gonad stages in Table 1. 1 = juvenile, 2 = 

 immature. 3 = active, 4 = developing, 5 = ripe. 6 = ripe, running, and 7 = spent. 



A positive relationship between temperature at the 

 time of spawning (back-calculated from stage-1 POFs, 

 assuming a delay of 6 hours) and the proportion of ova- 



ries with stage-1 POFs indicated that spawning events 

 were positively correlated with temperature (r=0.93) 

 over the range 9°C and 22°C (Fig. 9). GSI was how- 



