BURGESS: FOUR NEW SPECIES OF SQUID ENOPLOTEUTHIS 



Table 1.— Measurements (in millimeters) and counts of Enoploteuthis obliqua. 



missing mantle or arm tip, or lost suckers. 



TC = RV Townsend Cromwell; + = a 



'From stomach of Alepisaurus. 



of the neck that are connected to each other 

 posteriorly by a narrow membranous ridge. The 

 third dorsal fold extends dorsally as a mem- 

 branous ridge, but the ridge does not reach the 

 midline of the head. Dorsal and ventral ocular 

 "windows" are present and the latter easily allows 

 a count of the nine eye photophores on the ventral 

 side of the eyeball. The eye opening is a large, 

 wide, transverse oval with a deep sinus (Fig. 

 1 F). 



The buccal membrane completely hides the 

 buccal mass and surrounding lips. Eight slender 

 supports are joined by slender connectives to the 

 arms in the order DDVD (i.e., all are attached to 

 the dorsal side of the arms except the third pair 

 which are attached to the ventral side of arm III). 

 The inner surface of the buccal membrane is 

 rugose, but without papillae; the lappets are 

 delicate and pointed. The membrane is purple 

 and does not appear much darker than most 

 parts of the body. The chromatophores are small 

 and are scattered evenly on both the supports 

 and the membrane. 



The arms are subequal, slender, and much 

 shorter than the mantle length (ALI: I, 48.8- 

 50.2-52.0; II, 43.6-50.2-54.0; III, 47.3-5L0-6O.O; 

 IV, 48.8-55.2-62.0). They taper gradually to 

 slender tips. The swimming keels are low and 

 poorly developed on the dorsal arms, and they 

 are mainly confined to the distal third of these 



arms. The swimming keel of arm III is wider 

 than the arm at its greatest width. The lateral 

 membrane or tentacular sheath along arm IV is 

 narrow; it is about half of the arm width proxi- 

 mally and it extends to the tip of the arm. Protec- 

 tive membranes are developed on the ventral 

 side of all arms, but decrease in size in the follow- 

 ing order: III, II, I, IV. The dorsal protective 

 membranes on all the arms are low; on arm IV 

 the trabeculae are not evident. 



The right ventral arm of the male is hectocoty- 

 lized. The protective membrane on the medial 

 side of this arm (Fig. 1J) is expanded into an un- 

 dulating membrane that extends from the 

 eighth pair of hooks to the tip of the arm. The dor- 

 sal protective membrane is slightly developed. 

 In the males small tubercles or conical papillae 

 are present between the bases of the hooks and 

 bases of all arms. 



All the arms bear two rows of alternating 

 strong hooks, each (Fig. IE) completely enclosed 

 in a membranous sheath. The distalmost hook is 

 about half the length of the largest one. In both 

 sexes, arm IV has the most hooks. The distal part 

 of each arm is occupied by two rows of suckers 

 with wide apertures and slender stalks. The 

 inner sucker ring bears seven or eight prominent 

 truncated teeth on the distal margin but the 

 proximal margin is smooth (Fig. IB). The outer 

 ring has numerous pegs (Nixon and Dilly 1977). 



705 



