BURGESS: FOUR NKW SPECIES OF SQUID F.NOI'LOTEVTHIS 



130° 140° 150° 160° "170° 180° 170° 150° 150° 140° 130° 120° 110° 100° 90° 80° 70° 



Figure 7.— Distribution of Enoploteuthis species in the Pacific. 



higginsi, and reticulata. Most of the specimens 

 (93.1%) were taken between the surface and 150 

 m, and the greatest number of these (48. 1%) came 

 from a depth of 50 m. A majority of the 

 specimens (70.2%) were collected at night 

 between 1900 and 0400. A limited number of 

 specimens (8.4%) came from the few tows made 

 during the day. Enoploteuthis higginsi and E. 

 reticulata captured off Oahu, Hawaii, were 

 taken close to the surface (50-100 m) at night and 

 much deeper (500-600 m) during the day (Young 



1978). 



Species of Enoploteuthis are undoubtedly 

 mesopelagic forms (200-1,000 m day habitat) 

 that make diel vertical migrations as has been 

 demonstrated for E. a napsis by Roper ( 1966) and 

 for two other Enoploteuthis by Young (1978). 



RELATIONSHIPS 



The Pacific species of Enoploteuthis fall into 



two natural groups allied either to E. anapsis or 

 E. leptura along the same characters that 

 separate these two Atlantic species (see Roper 

 1966). Apparently, there are two species com- 

 plexes within the genus: Species that tend 

 toward the development of larger clubs and long 

 and muscular tentacles, and those species that 

 exhibit the opposite trends. 



Additional features of the former complex are: 

 many suckers in four rows on the clubs, large 

 hooks, and oval to round carpal apparatus. These 

 are features common to galaxias, chuni, 

 theragrae, anapsis, jonesi, and higginsi. Within 

 this group, anapsis, jonesi, and higginsi all have 

 the semilunar membrane on the club while 

 galaxias and theragae are illustrated in the 

 literature without it. Both Berry (1918) and Taki 

 (1964) do not mention the semilunar membrane 

 in their species descriptions. I have seen the 

 holotype of galaxias, but I could not verify the 

 presence of the membrane because of the poor 



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