Chen et al,: Population structure of Homarus americanus in the Gulf of Maine 



255 



size-dependent inshore-offshore distribution (or a size- 

 dependent distribution that is related to depth or other 

 environmental variables). The two sampling programs 

 can greatly improve the quality and quantity of data col- 

 lected in a fishery-independent program, leading to the 

 improved assessment and management of fisheries. In 

 the case where only one survey program can be conduct- 

 ed (e.g., due to budget limitation or time constraints, or 

 both), we suggest that area-specific fishery landing data, 

 together with depth or other environmental variables 

 that influence fish distributions, be used in allocating 

 sampling efforts in a random stratified survey. 



Acknowledgments 



We would like to thank R. Tetrault of T&R Fish Inc. for 

 providing the vessels, Capt. Curtis Rice of the FV Robert 

 Michael and Capt. Samuel Galli of the FV Tara Lynn for 

 contributing to the project from the start. We also would 

 like to thank V. Manfredi, J. Brown, K. Stepanek, and 

 D. Grout for field assistance and data entry and prepara- 

 tion. L. Jacobson and J. Idoine of the Northeast Fisheries 

 Science Center provided the NMFS survey information, 

 which we appreciate greatly. The financial support of 

 this study was provided by the NMFS. Maine Depart- 

 ment of Marine Resources, Northeast Consortium, and 

 Maine Sea Grant Program. 



Literature cited 



ASMFC (Atlantic State Marine Fisheries Commission). 



2000. The American lobster stock assessment, 532 

 p. ASMFC, Washington, D.C. 

 Briggs, P. T. 



1985. Movement of the American lobster off the south 

 shore of Long Island, New York. New York Fish Game 

 32:20-25. 



Caddy, J. F. 



1975. Spatial model for an exploited shellfish popula- 

 tion, and its application to the Georges Bank scallop 

 fishery. J. Fish. Res. Board Can. 32:1305-1328. 

 Campbell, A. 



1983. Growth of tagged lobsters off Port Maitland, Homa- 

 rus americanus, in the Bay of Fundy. Can. J. Fish. 

 Aquat. Sci. 40:1667-1675. 

 Campbell, A., and D. S. Pezzack 



1986. Relative egg production and abundance of berried 

 lobsters, Homarus americanus, in the Bay of Fundy and 

 off Southweatern Nova Scotia. Can. J. Fish. Aquat. 

 Sci. 43:2190-2196. 



Chen, Y., G. Liggins, K, J. Graham, and S. J. Kennelly. 



1997. Modelling length-dependent offshore distribution 

 of redfish, Centroberyx affinis. Fish. Res. 29:39-54. 

 Cobb, J. S., D. Wang, D. B. Campbell, and P. Rooney. 



1989. Speed and direction of swimming by postlar- 

 vae of the American lobster. Trans. Am. Fish. Soc. 

 8:82-86. 



Cooper, R. A., R. A. Clifford, and C. D. Newell. 



1975. Seasonal abundance of the American lobster, Homa- 

 rus americanus, in the Boothbay region of Maine. Trans. 

 Am. Fish. Soc. 104:669-674 

 Cooper, R. A., and J. R. Uzmann. 



1980. Ecology of juvenile and adult Homarus. In The 

 biology and management of lobsters, vol. 2. (J. S. Cobb 

 and R. Phillips, eds. I, p. 97-141. Academic Press, New 

 York, NY. 

 Harding, G. C, and R. W. Trites. 



1989. Dispersal of Homarus americanus larvae in the 

 Gulf of Maine from Brown Bank. Can. J. Fish. Aquat. 

 Sci. 46:1077-1078. 

 Hilborn, R., and C. Walters. 



1992. Quantitative fisheries stock assessment: choice, 

 dynamics, and uncertainty, 570 p. Chapman and Hall, 

 New York, NY. 

 Krouse, J. S. 



1980. Summary of lobster, Homarus americanus, tagging 

 studies in American waters (1898-1978). Can. Tech. 

 Rep. Fish. Aquat. Sci. 932:135-151. 

 Lawton, P., and K. L. Lavalli. 



1995. Postlarval, juvenile, adolescent and adult ecol- 

 ogy. In Biology of the lobster Homarus americanus 

 (J. F. Factor, ed.), p. 47-88. Academic Press, New 

 York, NY. 

 Palma, A. T, R. S. Steneck, and J. Wilson. 



1999. Settlement-driven, multiscale demographic patterns 

 of large benthic decapods in the Gulf of Maine. J. Exp. 

 Mar. Biol. Ecol. 241:107-136. 

 Paloheimo, J. E., and L. M. Dickie. 



1964. Abundance and fishing success. Rapp. P. V. Reun. 

 Cons. Int. Explor. Mer 1964:152-163. 

 Pezzack, D. S., and D. R. Duggan. 



1986. Evidence of migration and homing of lobsters 

 on the Scotian shelf Can. J. Fish. Aquat. Sci. 43: 

 2206-2211. 

 Rowling, K. 



1994. Tiger flathead, Neoplatycephalus richardsoni. In 

 The South East fishery (R. D. J. Tilzey, ed.), p. 

 124-136. Bureau of Resource Sciences, Australian 

 Gov. Print Serv., Canberra, Australia. 

 Skud, B. E. 



1969. The effect of fishing on size composition and sex 

 ratio of offshore lobster stocks. Fish Dir Skr. Ser. 

 HavUnders:15:295-309. 

 Taylor, C. C. 



1953. Nature of variability in trawl catches. Fish. Bull. 

 54:145-166. 

 Wahle, R. A. 



1992. Body-size dependent anti-predator mechanisms of 

 the American lobster. Oikos 65:52-60. 

 Wahle, R. A., and R. S. Steneck. 



1992. Habitat restrictions in early benthic life: experi- 

 ments on habitat selection and in situ predation 

 with the American lobster. J. Exp. Mar. Biol. Ecol. 

 157:91-114. 

 Wilson, C. J. 



1998. Bathymetric and spatial patterns of settlement in 

 American lobsters, H. americanus, in the Gulf of Maine: 

 Insights into processes controlling abundance. M.S. 

 thesis, 37 p. Univ. Maine, Orono, ME. 



