U.S. DEPARTMENT OF COMMERCE 



Malcolm Baldrige, Secretary 



NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



John V. Byrne, Administrator 



NATIONAL MARINE FISHERIES SERVICE 



William G. Gordon, Assistant Administrator 



Fishery Bulletin 



The Fishery Bulletin carries original research reports and technical notes on investigations in fishery science, engineering, and 

 economics. The Bulletin of the United States Fish Commission was begun in 1881; it became the Bulletin of the Bureau of Fisheries 

 in 1904 and the Fishery Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents through volume 46; the 

 last document was No. 1103. Beginning with volume 47 in 1931 and continuing through volume 62 in 1963, each separate appeared as 

 a numbered bulletin. A new system began in 1963 with volume 63 in which papers are bound together in a single issue of the bulletin 

 instead of being issued individually. Beginning with volume 70, number 1, January 1972, the Fishery Bulletin became a periodical, 

 issued quarterly. In this form, it is available by subscription from the Superintendent of Documents, U.S. Government Printing 

 Office, Washington, DC 20402. It is also available free in limited numbers to libraries, research institutions. State and Federal 

 agencies, and in exchange for other scientific publications. 



SCIENTIFIC EDITOR, Fishery Bulletin 



Dr. William J. Richards 



Southeast Fisheries Center Miami Laboratory 



National Marine Fisheries Service, NOAA 



Miami, FL 33149-1099 



Editorial Committee 



Dr. Bruce B. Collette Dr. Donald C. Malins 



National Marine Fisheries Service National Marine Fisheries Service 



Dr. Edward D. Houde Dr. Jerome J. Pella 



Chesapeake Biological Laboratory National Marine Fisheries Service 



Dr. Merton C. Ingham Dr. Jay C. Quast 



National Marine Fisheries Service National Marine Fisheries Service 



Dr. Reuben Lasker Dr. Carl J. Sindermann 



National Marine Fisheries Service National Marine Fisheries Service 



Mary S. Fukuyama, Managing Editor 



The Fishery Bulletin (ISSN 0090-06561 is published quarterly by the Scientific Publications Office, National Marine Fisheries 

 Service, NOAA, 7600 Sand Point Way NE, BIN C1570O, Seattle, WA 98115. Second class postage is paid at Seattle, Wash., and 

 additional offices. POSTMASTER send address changes for subscriptions to Superintendent of Documents, U.S. Government 

 Printing Office, Washington, DC 20402. 



Although the contents have not been copyrighted and may be reprinted entirely, reference to source is appreciated. 



The Secretary of Commerce has determined that the publication of this periodical is necessary in the transaction of the public 

 business required by law of this Department. Use of funds for printing of this periodical has been approved by the Director of the 

 Office of Management and Budget through 1 April 1985. 



