HAYNES: PANDALIDAE, HIPPOLYTIDAE. ANDCRANGONIDAE LARVAE 



For example, the spine is present in Stage I Pan- 

 dahis hessleri, but absent in P. hypsinotus until 

 Stage III. The stage at which the anal spine first ap- 

 pears in crangonid larvae with abbreviated develop- 

 ment has not been reported. 



KEY TO STAGE OF DEVELOPMENT 



Larvae of most of the species in this report can be 

 keyed to stage based on development of the eyes and 

 tail fan (Key I; Table 1). Key I is mostly applicable to 

 species whose larval development is not markedly 

 abbreviated (i.e., those with > 5 zoeal stages). In 

 species with unabbreviated development, exopodites 



on pereopods are characteristically undeveloped in 

 Stage I and usually have natatory setae beginning at 

 Stage II or III. Most of the species excluded from the 

 key (Table 2) have < 4 zoeal stages, and exopodites 

 on pereopods are either absent in all stages, vestigial 

 in Stages I and II, or have natatory setae as early as 

 Stage I. Key I is limited because it does not differen- 

 tiate between the latest stages and uses only one or 

 two characters, which may be absent in damaged 

 specimens. 



For pandalid shrimp. Key I can be supplemented 

 by keys to stages for each species (i.e.. Keys IV and 

 VI-XI, which have characters not given in Key I and 

 distinguish between the latest stages). With a few 



Table 1.— Species included In keys. 



Pandalidae Haworth 1825 



HIppolytidae Bate 1888 



Crangonidae White 1947 



Pandalopsis Bate 1888 

 P. cocclnata Urita 1941 

 P. d/spar Rathbun 1902 



Pandalus Leach 1814 

 P. boreal is Krdiyer 1838 

 P. danae Stimpson 1857 

 P. gon/urus Stimpson 1860 

 P. hypsinotus Brandt 1851 

 P. jordani Rathbun 1902 

 P. /(ess/en CzernlavskI 1878 

 P. platyceros Brandt 1851 

 P. prensor Stimpson 1860 

 P, stenolepis Rathbun 1902 

 P. tridens Rathbun 1902 



Eualus Thallwltz 1892 



E. barbatus (Rathbun 1899) 



£. fabricii (Kr6yer 1841) 



E. gaimardii (H. Milne Edwards 1837) 



E. herdmani (Walker 1898) 



E. macilentus (Krciyer 1841) 



£. pusiolus (Krdyer 1841) 



E. sucl<leyi (Stimpson 1864) 



Heptacarpus Holmes 1900 

 H. brevirostris (Dana 1852) 

 H. camtschaticus (Stimpson 1860) 

 H. paludicola (Holmes 1900) 

 H. tridens (Rathbun 1902) 



Hippolyte Leach 1815 

 H. clarki Chace 1951 



Lebbeus White 1847 



L. groenlandicus (Fabricius 1775) 

 L. Polaris (Sabine 1821) 



Spirontocaris Bate 1888 

 S. arcuata Rathbun 1902 

 S. /////etoo/'g// (Danlelssen 1859) 

 S. murdochi Rathbun 1902 

 S. ochotensis (Brandt 1851) 

 S. phippsii (Kr(4yer 1841) 

 S. spinas (Sowerby 1805) 

 S. spinas var. intermedia 

 Makarov 1941 



Argis Krciyer 1842 



A. crassa (Rathbun 1899) 

 A. dentata (Rathbun 1902) 

 A. lar {Owen 1839) 



Crangon Fabricius 1798 

 C. af finis de Haan 1849 

 C. alaskensis Locklngton 1877 

 C. communis Rathbun 1899 

 C. dalli Rathbun 1902 

 C. franciscorum angustimana 



Rathbun 1902 

 C. septemspinosa Say 1818 



Mesocrangon Zarenkov 1965 

 M. intermedia (Stimpson 1860) 



Paracrangon Dana 1852 

 P. echinata Dana 1852 



Sabinea J. C. Ross 1835 

 S. septemcarinata 

 (Sabine 1824) 



Sclerocrangon G. O. Sars 1883 

 S. boreas (Phipps 1774) 

 S. salebrosa (Owen 1839) 

 S. zenkevitchi Birstein and 

 Vinogradov 1953 



Table 2. — Species that cannot be keyed to stage of develop- 

 ment using Key I. 



Pandalidae 



Hippolytidae 



Crangonidae 



Pandalopsis 

 P. cocclnata 

 P. d I spar 

 Pandalus 

 P. danae 



(Stage III) 

 P. hypsinotus 



(Stage III) 

 P. kessleri 

 P. platyceros 

 P. prensor 

 P. stenolepis 



(Stage III) 



Lebbeus 



L. groenlandicus 

 L. polaris 



Argis 



A. crassa 



A. dentata 



A. lar 

 Sclerocrangon 



S. boreas 



S. salebrosa 



S. zenkevitchi 

 Sabinea 



S. septemcarinata 



263 



