U.S. DEPARTMENT OF COMMERCE 



Juanita M. Kreps, Secretary 



NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION 



Richard A. Frank, Administrator 



Terry L. Leitzell, Assistant Administrator for Fishieries 



NATIONAL MARINE FISHERIES SERVICE 



Fishery Bulletin 



The Fishery Bulletin carries original research reports and technical notes on investigations in fishery science, engineering, and 

 economics. The Bulletin ofthe United States Fish Commission was begun in 1881; it became the Bulletin of the Bureau of Fisheries in 

 1904 and the Fishery Bulletin ofthe Fish and Wildlife Service in 1941 . Separates were issued as documents through volume 46; the last 

 document was No. 1103. Beginning with volume 47 in 1931 and continuing through volume 62 in 1963, each separate appeared as a 

 numbered bulletin. A new system began in 1963 with volume 63 in which papers are bound together in a single is.sueof the bulletin 

 instead of being issued individually. Beginning with volume 70, number 1, January 1972, theFis/iery Bulletin became a periodical, 

 issued quarterly. In this form, it is available by subscription from the Superintendent of Documents, U.S. Government Printing Office, 

 Washington, DC 20402. It is also available free in limited numbers to libraries, research institutions. State and Federal agencies, and 

 in exchange for other scientific publications. 



EDITOR 



Dr. Jay C. Quast 



Scientific Editor. Fishery Bulletin 



Northwest and Ala.ska Fisheries Center 



Auke Bay Laboratory 



National Marine Fisheries Service, NOAA 



P.O. Box 155, Auke Bay, AK 99821 



Editorial Committee 



Dr. Elbert H. Ahlstrom 



National Marine Fisheries Service 



Dr. Bruce B. Collette 



National Marine Fisheries Service 



Dr. Edward D. Houde 

 University of Miami 



Dr. Merton C. Ingham 



National Marine Fisheries Service 



Dr. Reuben Lasker 



National Marine Fisheries Service 



Dr. Jerome J. Pella 



National Marine Fisheries Service 



Dr. Sally L. Richardson 



Gulf Coast Research Laboratory 



Kiyoshi G. Fukano, Managing Editor 



The F/she/y Bulletin (USPS 090-870) is published quarterly by Scientific Publications Office. National Marine Fisheries Service. 

 NOAA. Room 450. 1107 NE 45th Street. Seattle. WA 98105 Controlled circulation paid to Finance Department. USPS. 

 Washington, DC 20260. 



Although the contents have not been copyrighted and may be reprinted entirely, reference to source is appreciated 



The Secretary of Commerce has determined that the publication ot this periodical is necessary in the transaction of the public 

 business required by law of this Department Use of funds for printing of this periodical has been approved by the Director of the 

 Office of fyfanagement and Budget through 31 lylarch 1982, 



