650 



Fishery Bulletin 92(3), 1994 



Table 3 



Gonad weight and energy content of northern sand 

 lance, Ammodytes dubius, from Georges Bank. 



Males 



Females Total mean 



N 



Fork length (mm) 



Dry weight (gi 



Gonad 



i'< dry body 



weight) 

 Gonad (kcal/g) 



1 

 132 

 1.23 



7 

 12717.4 

 1.22+0.35 



14.33 26.36±2.72 



6.4 7.12±0.52 



8 

 12817.1 

 1.23+4.94 



24.85+4.94 

 7.0310.54 



ber to March, to 2.90 kcal in May (Table 2). The mean 

 percent of body weight accounted for by the ripe go- 

 nads, 24.85%, is similar to the 20-30% measured for 

 age 1+ A. personatus (Okamoto et al., 1989) and the 

 25-28% measured for both male and female A. 

 americanus (Smigielski et al., 1984), (Table 3). Esti- 

 mated monthly metabolic requirements range from 

 a low in February of 2.03 kcal to a high in May of 

 4.15 kcal (Table 4). The annual individual energy 

 requirement was 37.56 kcal (Table 4). Monthly as- 

 similation efficiencies and daily rations are shown 

 in Table 5, and the predicted annual ration is 52.62 

 kcalfish -1 or 8.60 g-fish -1 (assuming a caloric con- 

 tent of 6.11 kcalg -1 for C. finmarchicus; Larimer, 

 1992). The energy assimilated from that ingested is 

 48.09 kcal-fish -1 so the annual individual energy 

 budget in terms of kilocalories is approximated by 



(G) + (R) = (AR) -(/?), 



(3) 



where growth (G) = 8.08 kcal-yr -1 ; 



reproduction (R) = 2.45 kcal-yr -1 ; 



assimilated ration (AR) = 48.09 kcal-yr -1 ; 



respiration (R) = 37.56 kcal-yr -1 . 



If the budget is converted into percentage of total 

 consumption accounted for by each parameter, the 

 following relationship results: 



