U.S. Department 

 of Commerce 



Donaid L. Evans 

 Secretary 



National Oceanic 

 and Atmospheric 

 Administration 



Vice Admiral 



Conrad C. Lautenbacher Jr., 



USN (ret.) 



Under Secretary for 

 Oceans and Atmosphere 



National Marine 

 Fisheries Service 



William T. Hogarth 



Assistant Administrator 

 for Fisheries 



.^TOFCo. 



X 



K 1 ^ / 



The Fishery' Bulletin (ISSN 0090-0656) 

 is published quarterly by the Scientific 

 Publications Office, National Marine Fish- 

 N( >AA, 7600 Sand Point Way 

 NE, BIN C15700, Seattle. WA 981 15-0070. 

 Periodicals postage is paid at Seattle, WA, 

 and at additional mailing offices. POST- 

 MASTER: Send address changes for sub- 

 scription- to Fishery Bulletin. Superin- 

 tendent of Documents, Attn.: Chief. .Mail 

 List Branch, Mail Stop SSOM, Washing- 

 ton. DC 20402-9373. 



Although the contents of this publica- 

 tion have not been copyrighted and may 

 be reprinted entirely, reference to source 

 is appreciated. 



The Set if Commerce has deter- 



mined that the publication of tin 



ording to law for 



the transaction of public business of this 



Department. Use of funds for printing of 



nodical has been approved by the 



oroftheOffii cement and 



Budget. 



For sale by the Superintendent of 

 nuts. US. Government Printing 

 I mice, Washington, DC 20402. Subscrip- 

 tion pi i it: $55.00 domestic and 

 $68.75 foreign. Cost per single issue: 

 $28.00 dome ,5.00 foreign. See 

 back for order form. 



Scientific Editor 



Dr. Norman Bartoo 



Associate Editor 



Sarah Shoffler 



National Marine Fisheries Service, NOAA 

 8604 La Jolla Shores Drive 

 La Jolla, California 92037 



Managing Editor 



Sharyn Matriotti 



National Marine Fisheries Service 

 Scientific Publications Office 

 7600 Sand Point Way NE, BIN C15700 

 Seattle, Washington 981 15-0070 



Editorial Committee 



Dr. Harlyn O. Halvorson 

 Dr. Ronald W. Hardy 

 Dr. Richard D. Methot 

 Dr. Theodore W. Pietsch 

 Dr. Joseph E. Powers 

 Dr. Harald Rosenthal 

 Dr. Fredric M. Serchuk 

 Dr. George Watters 



University of Massachusetts, Boston 

 University of Idaho, Hagerman 

 National Marine Fisheries Service 

 University of Washington, Seattle 

 National Marine Fisheries Service 

 Universitat Kiel, Germany 

 National Marine Fisheries Service 

 National Marine Fisheries Service 



Fishery Bulletin web site: www.fishbull.noaa.gov 



The Fishery Bulletin carries original research reports and technical notes on investigations in 

 fishery scien ring, and economics. It began as the Bulletin of the United States Pish 



Commission in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery 

 Bulletin of the Fish and Wildlife Service in 1941. Separates were issued as documents tl 

 volume 46; the last document was No. 1103. Beginning with volume 47 in 1931 and continuing 

 through volume 62 in 196.1 ired as a numbered bulletin. A new system 



began in 1963 with volume 6:3 in which papers are bound together in a single issue of the 

 bulletin. Beginning with volume 70. number 1. January 1972, the Fishery Bulletin became a 

 lieal, issued quarterly. In this form, it is available by subscription from the Superintendent 

 of Documents. U.S. Government Printing Office, Washington, DC 20402. It is also available free in 

 limited numbers to libl irch institutions. State and Federal agencies, and in exi 



for other scientific publications. 



