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327 



is the number of removals of the cohort due to natural 

 mortality up to time t (Quinn and Deriso, 1999) and E 

 (when no fishing takes place): 



E=\ E SP jdD t = f E^( 1 - e" 1 " )dD l 



= EpN \(l-e- l ")e' m 'mdt, 

 o 



where D t = N (l-e- mt ) is the number of removals of the 

 cohort due to natural mortality when no fishing takes 

 place. 



