U.S. Department 

 of Commerce 



Donald L. Evans 

 Secretary 



National Oceanic 

 and Atmospheric 

 Administration 



Vice Admiral 



Conrad C. Lautenbacher Jr., 



USN (ret.) 



Under Secretary for 

 Oceans and Atmosphere 



National Marine 

 Fisheries Service 



William T. Hogarth 



Assistant Administrator 

 for Fisheries 



^tof c 



**r E s of * 



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Scientific Editor 



Norman Bartoo, PhD 



Associate Editor 



Sarah Shoffler 



National Marine Fisheries Service, NOAA 

 8604 La Jolla Shores Drive 

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Managing Editor 



Sharyn Matriotti 



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Editorial Committee 



Harlyn O. Halvorson, PhD 

 Ronald W. Hardy, PhD 

 Richard D. Methot, PhD 

 Theodore W. Pietsch, PhD 

 Joseph E. Powers, PhD 

 Harald Rosenthal, PhD 

 Fredric M. Serchuk, PhD 

 George Watters, PhD 



University of Massachusetts, Boston 

 University of Idaho, Hagerman 

 National Marine Fisheries Service 

 University of Washington, Seattle 

 National Marine Fisheries Service 

 Universitat Kiel, Germany 

 National Marine Fisheries Service 

 National Marine Fisheries Service 



Fishery Bulletin web site: www.fishbull.noaa.gov 



The /•' iid technical not ions in 



[I began as the Bulletin of the United Stai 

 Commission in 1881; it became the Bulletin of the Bureau of Fisheries m 1904 and the I 

 Bulletin of the Fish and Wildlife S 941. Separates were issued as documents tl 



volume Hi. the last document was No. 1103. Beginning with volume 47 in 1931 and continuing 



Ii volume 82 in IS a numbered bulletin. A new system 



i which papers are bound together in a single issue of the 

 bulletii f0, number 1. January 1972. the Fishery Bulletin bee: 



periodical, - terly. In this form, it is available by subscription from the Superintendent 



uments, U.S. Gove nt Printing Office, Washington. DC 20402. Ii i- also available free in 



limited numbers to hi" arch institutions, Slate and Federal agencies, and in exchange 



lor other scientific publications 



