472 



Fishery Bulletin 102(3) 



hunters) in all cells, 3) single tag recoveries by fishermen 

 in all cells, and 4) the data set employed in the present 

 study of single-tag recoveries by fishermen in one of two 

 cells, the latter represents the low end in quality and 

 quantity of information about movement and survival. 



A time-dependent approach could theoretically extend 

 the approach of McGarvey and Feenstra (2002) to make 

 explicit the residence times of each recaptured indi- 

 vidual in the fishing zone and sanctuary, respectively, 

 and thus make explicit differences in the predicted 

 survival rate before and after movement. However with- 

 out prior knowledge of when movement took place for 

 each recaptured lobster, a modified likelihood method 

 is called for, requiring integration over the probable 

 movement times between tag release and recapture. 

 This extension of residence-time-dependent movement 

 estimators to variable times of movement remains a 

 topic for future research. 



Acknowledgments 



I thank Hugh Possingham, Andre Punt, and two review- 

 ers for comments on the draft manuscript. Lobsters 

 were tagged and released into Gleesons Landing sanctu- 

 ary by Greg Ferguson, together with fishermen Lenny 

 and Murray Williams, under Fisheries Research and 

 Development Corporation Project 93/086. This work was 

 supported by the Australian Fisheries Research and 

 Development Corporation Project No. 2000/195, and by 

 the South Australian rock lobster industry. 



Literature cited 



Abramowitz, M.. and I. R. Stegun. 



1965. Handbook of mathematical functions, p. 257. Dover 

 Publications, New York, NY. 

 Arnason, A. N. 



1972. Parameter estimates from mark-recapture exper- 

 iments on two populations subject to migration and 

 death. Res. Popul. Ecol. (Kyoto) 13:97-113. 



1973. The estimation of population size, migration rates, 

 and survival in a stratified population. Res. Popul. 

 Ecol. (Kyoto) 15:1-8. 



Booth, J. D. 



1997. Long-distance movements in Jasus spp. and their 

 role in larval recruitment. B. Mar. Sci. 61ill: 1 11— 

 128. 

 Brownie, C, J. E. Hines. J. D. Nichols, K. H. Pollock, and 

 J. B. Hestbeck. 



1993. Capture-recapture studies for multiple strata in- 

 cluding non-Markovian transitions. Biometrics 49:1173- 

 1187. 

 Gardner, C, S. Frusher, M. Haddon, and C. Buxton. 



2003. Movements of the southern rock lobster Jasus 

 edwardsii in Tasmania, Australia. Bull. Mar. Sci. 73(3): 

 653-671. 

 Hestbeck, J. B. 



1995. Bias in transition-specific survival and movement 

 probabilities estimated using capture-recapture data. J. 

 Appl. Stat. 22:737-750. 

 Hilborn, R. 



1990. Determination offish movement patterns from tag 

 recoveries using maximum likelihood estimators. Can. 

 J. Fish. Aquat. Sci. 47:635-643. 



Kelly, S. 



2001. Temporal variation in the movement of the 

 spiny lobster Jasus edwardsii. Mar. Freshw. Res. 

 53(31:323-331. 



Kelly, S., and A. B. MacDiarmid. 



2003. Movement patterns of mature spiny lobsters, Jasus 



edwardsii, from a marine reserve. N.Z. J. Mar. Freshw. 



Res. 37:149-158. 

 MacDiarmid, A. B.. B. Hickey, and R. A. Mailer. 



1991. Daily movement patterns of the spiny lobster Jasus 

 edwardsii (Hutton) on a shallow reef in northern New 

 Zealand. J. Exp. Mar. Biol. Ecol. 147:185-205. 



McGarvey, R., and J. E. Feenstra. 



2002. Estimating rates of fish movement from tag recov- 

 eries: conditioning by recapture. Can. J. Fish. Aquat. 

 Sci. 59:1054-1064. 



Quinn, T J. II, and R. B. Deriso. 



1999. Quantitative fish dynamics, p. 414-419. Oxford 

 Univ. Press, New York. NY. 

 Rice, J. A. 



1995. Mathematical statistics and data analysis, p. 13-14, 

 39-40. Duxbury Press, Belmont CA. 

 Schwarz, C. J., J. F. Schweigert, and A. N. Arnason. 



1993. Estimating migration rates using tag recovery 

 data. Biometrics 49:177-193. 

 Seber, G. A. F. 



1982. The estimation of animal abundance and related 

 parameters, 2nd ed., p. 59-60. Griffin, London. 



