368 Transactions. — Botany. 



C. monroi, Hooker,* and C. nana, Colenso.t 



Both of these differ in habit from C. flagelliformis. Their 

 growth resembles that of alpine plants. G. nana is a "very 

 dwarfy, glabrous, rigid shrub, 2 in. -4 in. high, with much- 

 compressed minutely striated branchlets Jg- in. -i in. in 

 diameter" (Hooker). G. monroi grows higher, reaching about 

 1 ft. in height, but much more closely, forming a very firm 

 cushion of circular form, in which, however, the individual 

 branches and branchlets are easily separated and dis- 

 tinguished. In neither is there any sign of leaves in the 

 adult form, and they flower later than Garmicliaclia flagel- 

 liformis. The tips of the shoots in G. monroi are coloured 

 brownish-red. 



Stem- structure.- — In general stem-structure they resemble 

 G. flagelliformis, but the chlorophyll tissue is developed in 

 greater abundance in proportion to the stereom tissue than 

 in the latter. There is the same double-layered epidermis 

 with thick external cuticle ; in G. monroi there are slight 

 ridges above the stereom strands, while the stomata occur in 

 the depressions, underneath which is the chlorophyll tissue. 

 Underneath each stoma is an air-cavity. The chlorenchyma 

 tissue extends inwards as far as the vascular bundles, and is 

 of palisade type in both G. monroi and G. nana, without any 

 air-spaces even in the inner layers. (Compare G. flagelli- 

 formis, where air-spaces are present.) 



In C. monroi the stereom tissue may or may not extend 

 inwards as far as the vascular bundle (fig. 9) ; when it does 

 not there are occasional small isolated groups alternating with 

 the peripheral groups, and occurring in the chlorophyll tissue. 

 Both these groups have a layer of polygonal thin-walled cells 

 around them. Where the stereom tissue does not extend as 

 far as the vascular bundle there is a crescent-shaped group of 

 stereom cells on the outside of the phloem, with a half-circle 

 of thin-walled cells round the outside ; there are also a few 

 similar thick-walled cells on the inside of the xylem, but with 

 no special thin- walled cells round them. The pith is com- 

 posed of large polygonal cells fitting together, without inter- 

 cellular spaces. The medullary-ray cells are similar in shape, 

 but somewhat smaller ; they pass over gradually into the 

 chlorenchyma cells. The stereom cells are polygonal, tightly 

 packed together ; when the shoot is quite young, as also in 

 G. flagelliformis, the cell-walls are quite thin, and the stem 

 is then very soft ; but it soon loses its softness and becomes 

 firm owing to the thickening of the cell-walls. 



* Hab. South Island. 



f Hab. North Island : Dry mountainous country at base of Mount 

 Tongariro. South Island: Southern Alps ; Otago, Waitaki River. 



