352 



Transactions . 



The vascular system is well developed. I found no bundle-sheath 

 present. 



The following references show that the structure of the so-called cortex 

 is constant throughout the genus :■ — 



Ganong (3, b) describes Salicornia herbacea as having " a branching, 

 succulent, practically leafless stem . . . possessing a compact stele 

 (with cortical system of bundles replacing those of abandoned leaves), thick 



water-storing cortex. 



Warming (2) says that in Salicornia ambigua the leaves stand out like 

 collars round the older parts of the branches, which are shrivelled and 

 thin, and in structure it differs little from Salicornia herbacea, described by 

 De Bary. 



De Bary (1, b) says Salicornia herbacea has chlorophyll tissue in pali- 

 sade form in the cortex of the stem, and has short scaly leaves arranged 

 in decussate pairs. 



Reasons for calling Leaf-base what appears to he and has been descnbed as 



" Cortex ." 



1. The vascular-bundle system in the cortex-like portion resembles 

 that of a leaf. The leaf-trace divides, the middle branch behaving nor- 

 mally : the two lateral branches are directed upwards for a short distance. 



Pa Hsade tissue 

 Storm / Aqueous 



~(~7\ tissue 



Oennatoge) 



Penblem 



'Plervwe 



Ftg. 3. — a, Longitudinal section of growing-point; b, c, x and y of fig. 2 



under higher power. 



and then run downwards in the aqueous tissue, anastomosing freely, and 

 forming a network. This position of the vascular bundles is comparable 

 to that in the basal portion of peltate leaves. 



2. Except for the median branch, the network has no connection with 

 the stem. 

 |g| 3. The bundles end blindly in mesophyll. 



? ! 4. There is no difference between the palisade-cells in the leaf and leaf- 

 base, and there is no break in continuity, they being developed in both 

 on the ventral side. 



