CONTENTS 



Page 



1. Introduction 1 



2. Formulae in SKEBUB 3 



Monthly changes in biomass 3 



Apex predators 3 



Fish species and benthos 3 



Zooplankton and phytoplankton ^ 



Components of the biomass equation k 



Growth (GS|^ ^) ^ 



Natural mortality (GMS ,) 5 



Consumption by apex predators (SS) 6 



Consumption by other species (CCj^ ^) 7 



Total food requirement of each species grouping (FOODj^) 7 



Amount consumed of each prey by each predator (CPN|^ 1^) 8 



Calculation of equilibrium biomasses 9 



3. Simulation of food suitability and availability dependent feeding 10 



Simulations of feeding in SKEBUB 12 



Resul ts and Limi tat ions 13 



h. Preliminary results from SKEBUB 13 



5 . References 1° 



6. Appendix tables 25 



