Table 5. t/pper; Transitive Closure Matrix for the Figure 1 Model of Total 

 Influence, as Summed Daily Carbon Fractions of Column Com- 

 partments, Transferred to Row Compartments Over all Paths of 

 all Lengths. 



Middle: Total Indirect Influence, as Summed Daily Carbon 

 Fractionsto Row Compartments Over all Pathsof all Lengths>1 . 

 Lower: Ratios of Total Indirect Influence (Middle) to Direct 

 Influence (Table 1 , Lower) from Column to Row Compartments 

 (°c Denotes Division by 0, i Denotes Indeterminate, 0/0). 



different compartments. For example, in its relationships to food source compart- 

 ments, pelagic carnivores (3) have indirect direct carbon flow ratios ranging from 1 3 

 to 1880 (row 3. lower Table 5 matrix), it is doubtful under these circumstances that 

 the pelagic carnivore populations could meaningfully adapt to their prey populations 

 based only on dietary composition of the latter. It is doubtful from the Table 5 figures 

 in general how adaptation of any kind could be possible in the Figure I system. And it 

 seems even more absurd to think that adaptation could occur in real, temporally 



101 



