628 THE EOCENE FAUNA. 



There are eight distinct carpal bones, those of the second series alter- 

 nating with those of the first, and also with the metacarpals. The first bone 

 of the second row, pi-obably the trapezium, was found displaced to a posi- 

 tion behind the others. It carries no facet for a first metacarpal. There 

 are four well-developed metacarpals, of which the V is shorter than the II. 

 They carry the usual keel on the posterior distal face. The phalanges are 

 rather short, and the unguis of digit III is w^ell expanded. 



The femur resembles in general the one figured by Owen as that of 

 Pliolophus vuJpiceps* and that of P. vintamisf figured by myself The 

 great trochanter projects far above the head, and there is ver}- little neck 

 The fossa Ugamenti teris is large, as is the trochanteric fossa. The third 

 trochanter is large, but less produced than in Tapirus and Equus. The 

 rotular crests are about equally elevated, thus resembling Tapirus more 

 than other recent Perissodactyla. The tibia is long and slender. It has a 

 prominent crest, which is cut off" from the external border of the head by a 

 deep notch. The crest is rather more prominent than in recent genera. 

 There are both proximal and distal faces of attachment for the fibula, but 

 I do not find the bone among those preserved. The inner malleolus is 

 short, and its external face is obliquely beveled in front. It carries no 

 facets, and is marked by a wade open vertical groove behind the middle. 

 The trochlear gx'ooves are deep and quite oblique The patella is robust. 



The calcaneum is elongate and compressed. The astragalar trochlea 

 is deeply grooved and has subvertical sides. The head is short but not 

 sessile, and carries two distal facets. The external is quite narrow and fits 

 the cuboid bone. The latter is longer than deep and deeper than wide. 

 The peroneal hook is present. The navicular is relatively deeper than in 

 the horse, and is closely united to the ecto- and mesocuneiform. The ento- 

 cuneiform is a large flat and oval bone, on the posterior side of the tarsus. 

 The mesocuneiform is the smallest. There are only thi-ee metatarsal bones, 

 and no laidiment of a fourth. Each one is applied to the entire distal sur- 

 face of its corresponding cuneifoi-m bone, without oblique articulation with 

 an adjoining one. The distal articular surface of the median metatarsal has 



* Quarterly Jouin. Geol. Soc, London, xiv, p. 54. 



t Report U. S. Geol. Geog. Expl. Snrr. W. of 100th Mer., G. M. Wheeler, iv, pi. Ixv. 



