BY J, J. FliETCHER. 483 



tively insignificant, while on legs in front of the seventh pair neither 

 papillae nor pores are visible ; in one specimen the legs of several 

 pairs have each two papillae, one above the other ; sometimes the 

 appearance of a papilla is exaggerated by the proximal portion of 

 the duct of the crural gland being slightly everted ; but I have 

 never yet seen a specimen in which papillae were present only on 

 the legs of the last pair. Not all the legs even of the same speci- 

 men are equally favourable for examination, but in well preserved 

 specimens even when papillae are not recognisable the apertures of 

 the ducts of the crural glands sometimes are. 



The papillse are round, usually whitish but sometimes not 

 different in colour from the surroundings, slightly post-axial in 

 position, and are situated near the base of the leg slightly external 

 to the nephridiopore. Mr. Sedgwick says : " It (the papilla) is in 

 the same position with regard to the leg as the corresponding 

 structure in the Cape males." (Monograph, Q.J.M.S. XXVIII, 

 p. 464), i. e., " on the second row of papillae counting from the 

 innermost pad " (1. c. p. 448). In the specimens examined by 

 me the papillie are located nearer the base of the leg than this, 

 on about the fifth-seventh row of papillse above the innermost 

 pad {i.e. about the third or fourth ridge below the nephridiopore); 

 I have never seen them so close to the pad as the second row. 



As was to be anticipated sections showed crural glands to be 

 present ; and as in P. capensis* each crural gland consists of a 

 dilated vesicular portion placed in the lateral compartment of the 

 body cavity in the leg, and of a narrow duct opening to the 

 exterior, on one of the papillse in question. As some of the 

 specimens show papillae on all the legs but those of the first pair, 

 it is a reasonable inference that in the males of P. leuckarti a 

 crural gland may normally be present in each leg but those of the 

 first pair. On the other hand the presence of two papillse on 

 some of the individual legs in one specimen, together with the 

 occasional absence of both papillse and pores [as in the last pair 

 of legs of each of two males of which sections were cut] shows 



* Memorial Edition of Balfour's Works, Vol. I. p. 905. 



