298 University of California Puhlications in Botany [Vol. 6 



carp irregular, due to infolding of wall but without projections ; inner 

 surface of wall verrucose, verrucosities narrower and lower than those 

 of outer ; primary cortex of large-celled pseudoparenchymatous tissue 

 to depth of verrucosities, below which tissue abruptly changes to 

 hyphal ; asci cylindrical, constricted between spores, rounded at apex, 

 30 by 240^ ; spores 1-seriate, ellipsoid, 24-28 by 32-34;u., surface cov- 

 ered with large, irregular, more or less densely crowded, flattened or 

 conical papillae rarely reaching 4/x in height and Ifx in width (gen- 

 erally 2 by 4) ; spores 1-seriate, placed irregularly in ascus; para- 

 physes delicate, 3-8/x wide (generally 4/x) arising from hyphal tissue 

 of primary cortex, and broadening above asci to form secondary 

 cortex. 

 "Found in forest, Mt. Tamalpais, Marin Co., Calif., April." 



No. 86, Harkness Collection, type. 



While the spore surface of this species is similar to that illustrated 

 by Tulasne (Fungi Hypogaei, tab. XIII) for G. Klotzschii, and answers 

 to the description by Fischer of the type specimen of Berkeley and 

 Broome (1897&, p. 24), the spore and ascus measurements are too 

 small, those given for G. Klotzschii being for the former 31-45 by 

 21-32^ and for the latter, 270-320 by 35-42/x. Also, the color of the 

 Californian species is lighter, it does not have the decided folds from 

 the base to the apex described by Fischer as usually present, while the 

 hyphal character of the primary cortex which is very marked in this 

 species has been found in no description of G. Klotzschii. In the her- 

 barium of the University of California is an Italian specimen from 

 Mattirolo labeled G. Klotzschii which differs from our species in a 

 very much more minutely verrucose surface, darker color of ascocarp, 

 the presence of occasional strands of sterile tissue in primary cortex, 

 larger and more nearly globular spores, and in the sculpturing of the 

 spore surface, which in this case consists of very irregular papillae, 

 the larger usually truncate and measuring sometimes 7 by Sp.. The 

 specimen from Mattirolo represents the variation from the type plant 

 cited by Fischer from Mattirolo 's herbarium (18976, p. 24), but the 

 notes would indicate that the variation is only in the spore and the 

 differences in ascocarp characters therefore would separate our plant 

 as a distinct species. 



The spore is very similar to Tulasne 's illustration of the spore of 

 G. hispid ula, and the spore and ascus measurements agree with those 

 cited for this species, but the lack of hispid character of the ascocarp, 

 the lighter brown color, irregular cavity and presence of hyphal tissue 

 (which is not described for G. hispidula and, from Tulasne 's illustra- 

 tion, is evidently not present) would separate the two species. 



