18 University of Calif ornia Puhlications in Botany [Vol. 6 



3. "Janczewskia australis? Falkbg." in Eeinbold, Meeres 



Algen von Investigator Street (Siid Australien). Hedwigia, vol. 38, p. 47 

 1899; De Toni, Syll. Alg., vol. 4, sect. 3, p. 812, 1903 (under /. tasma- 

 nica with a query). 



Apparently this nomen nudum was intended for J. tasmanica to 

 apply to a plant found on a Laurencia, determined as L. ohtusa, sent 

 to Reinbold from Investigator Strait near Adelaide. South Australia. 

 In view of the results of the investigations on California species de- 

 tailed above, it seems that further investigation of this plant is needed 

 to settle definitely its exact status within the genus. It seems probable 

 to the writer that this plant may belong to a distinct and, as yet, 

 unnamed species, since Reinbold compares it with the figure in the 

 Nati'uiichen Pflanzenfamilien (I. 2, p. 431, f. 243 C). which Falken- 

 berg later (1901, p. 257) states does not properly represent his Janc- 

 zewskia tasmanica. 



VII. RELATIONSHIPS 



In the first place are to be considered the relationships between 

 the genus Janczeivskia and other genera. These have been adequately 

 dealt vnth by Solms-Laubach (1877, p. 217). who recognized his plant 

 as a member of the Rhodomelaceae and not distantly removed from 

 Laurencia itself. Schmitz and Falkenberg (1897, p. 432) and all 

 following writers have placed Janczeivskia in the Rhodomelaceae in 

 close proximity to Laurencia in the sul3-family Laurencieae. Schmitz 

 (1893, p. 390) has called attention to the fact that Janczeivskia is one 

 of the several parasitic Florideae whose host plant is a member of the 

 same famil.y, and Batters (1892, p. 66). in describing Gonimopliyllum 

 Buffhami, a delesseriaceous parasite on the delesseriaceous genus 

 Nitophyllum, also speaks of Actinococcus and Janczewskia as examples 

 of Florideae parasitic on plants of the same family to which they 

 themselves belong. All four new species proposed above have cysto- 

 carps which agree in all essential details M'ith those of Janczewskia 

 verrucaeformis and with those of the Rhodomelaceae in general. Be- 

 sides the structure of the cj'stocarp, all the species of Janczewskia, 

 including the four new species described above, have the same sort of 

 apical pit with the apical cell projecting slightly from the bottom of 

 it, which is characteristic of the Laurencieae and not found in any 

 of the other sub-families, although something remotely like it occurs 

 among the Chondrieae. 



