1918] Crarcbier: New Pacific Coast Marine Algae II 431 



observed in two of the species, nor liave the characteristic basal fila- 

 ments been seen in any of them. More will be said concerning these 

 under each species. In placing the species described in this paper 

 the genus Hyella is being interpreted as having basal filaments which 

 give rise to erect filaments penetrating the host, and as having goni- 

 dangia formed from special cells just within the host, or formed by 

 the transformation of certain basal, surface cells at the basal ends 

 of the erect filaments. It is also assumed that these conditions prevail 

 in the species which are here presented although they have not been 

 observed, judgment being based upon the resemblance of the plants 

 as a whole to the original H. caespitosa. 



Sauvageau (1895) founded the genus Radaisia on R. Gomontmna. 

 He does not mention the basal layer or any of the early stages of 

 development, either in the diagnosis or in the appended notes. The 

 plant is represented as being composed of a mass of erect, nearly 

 parallel rows of spherical cells or filaments embedded in a coherent 

 jelly. The filaments are practically independent and are sparsely 

 branched, thus producing a fan-shaped appearance in longitudinal 

 section view. The gonidangia are terminal or intercalary, and the 

 gonidia are formed bv successive divisions of the cell contents. He 

 separates this genus from Entophysalis because of the filamentous 

 character of the cells in the colony, and particularly because of the 

 presence of gonidangia. He places his Entophysalis Cornu-ana in 

 Radaisia on account of its filamentous nature although gonidangia 

 had not been observed. 



Nothing being known of the method of the origin of a colony of 

 R. Goniontiana or of R. Cornuana, one must remain somewhat in 

 doubt when placing a species, whose early stages of development are 

 known, in Radaisia. There are at least two possibilities as to the 

 origin of the masses of perpendicular filaments of R. Gomontiana and 

 R. Cornuana. Tlie gonidia may escape in mass, come to rest on a 

 substratum, and then spr(>ad out. Later, these gonidial cells by divi- 

 sions in one horizontal plane would produce a new colony, each cell 

 giving rise to a single chain of cells, or a filament. The colony would 

 thus consist of a group of practically' independent plants. Or on 

 the contrary, the gonidia may escape and come to rest singly on the 

 substratum, and each one after locating may produce a prostrate, 

 or basal layer of cells by divisions in two or more planes, then by 

 subsequent horizontal divisions of these cells produce the erect fila- 

 ments. In the first case, the colony would consist of a number of 



