10 University of California Publications in Botany [Vol. g 



is no considerable difference between the cells of the tubercle and 

 those of the host as to size, shape, walls, or contents, so that it is not 

 an easy matter to decide just where the one ceases and the other begins. 

 In young plants it is possible to find the apical pits of the coalescent 

 branchlets thickly scattered through the surface tissues, but in adult 

 plants each apical pit shows reproductive bodies and have conse- 

 quently suffered change. The tetrasporangia are subepidermal, tripar- 

 tite or cruciate, and line the walls of conceptacles, or enlarged apical 

 pits, which thickly stud the outer surface of the tubercle and which 

 bear leaves (pi. 3, fig. 18; pi. 5, fig. 26). The antheridia are in broad, 

 ovate-pyriform conceptacles, arising in a more or less conical mass 

 from its broad, slightly rounded bottom (pi. 3, fig. 19; pi. 5, fig. 27). 

 Each tuft of antheridia is broadly conical (broadly triangular as seen 

 in radial section) and the main axis is almost equaled by the several 

 oblique lateral axes from towards the base of the cluster, making a 

 bouquet-like mass. These tufts resemble those figured for Janczewskia 

 verrucaeformis by Solms-Laubach (1877, pi. 3, fig. 15), but they do 

 not end in moniliformly swollen terminal hairs of a deep yellow color. 

 The cystocarpic plants are thickly covered with nearly glo])ular cysto- 

 carps in various stages of emergence from the frond (pi. 2, fig. 7; 

 pi. 3, fig. 17). They may be compared, also, to Solms-Laubach 's 

 figures (1877, pi. 3, figs. 7, 17) of the cystocarps of J. verrucaeformis, 

 which they resemble, but the pericarp ol" the adult cystocarps is as 

 thick ;is, or nearly as thick as, tlic horizontal diameter of the cavity, 

 consequently much thicker proportionally than as figured for J. ver- 

 rucaeformis. The carpostome seems to l)e a narrow, elongated aperture 

 at the apex of the pericar[). wliicli is not contracted just below it. 



Janczewskia Solmsii resembles closely J. verrucaeformis, and these 

 two species are undoubtedly nnich more closely related to one another 

 1 liaii any other two of the six species to be considered here. J. Solmsii, 

 however, diff'ei-s from Ihe descriptions and figures of J. verrucaeformis 

 ill being somewhat larger, in lacking color, having a different dispo- 

 sition (>r till" tetrasporangia, having a somewhat differently shaped, 

 thicker-walled cystocarp, and in lacking the apparently conspicuous 

 slci'ilc, iiioiiilifoi'inly swollen hairs on tlie antheridia! lufts. Adding 

 to llicsc ( I i ff'(!i'ences a difTcrcnt species of Lnurencia as a host and a 

 witlely separated locality, it seems best to consider it distinct and to 

 describe^ if as a sc^parate species, which Mr. Guernsey and the writer 

 take pleasui-e in dedicating to Professor Hermann Craf zu Solms- 

 Laubach. the discoverer of the lypc sjx'cics and foundci' of fh(> genus. 



