14 American Midland Naturalist Monograph No. 1 



Argas REFLEXUS (Fabricius), 1794 



Plate 1, Figs. 2 and 3 



1793. Acarus columbarum Shaw, in Shaw and Nodder, original description, 4, pi. 128. 



(nomen nudum). 

 1794 Acarus reflexus Fabricius, p. 426. 

 1802. Argas reflexus (Fabricius) : Latreille, p. 66. 



1804. Rhpnchoprion columbae Hermann, p. 69. 



1805. Ixodes reflexus and Ixodes colmbae Fabricius, p. 353 and 356. 



1815. Argas marginaius Oken, p. 402. 



1816. Rhvmchoprion marginatum Olfers, p. 75. 



1827. Ixodes hispanus and Ixodes espagnol. Brebisson, p. 267. 



1828. Acarus marginatus. Guerin-Meneville, p. 300. 



1829. Argas reflexus. Latreille, p. 289. 



1896. Argas magnus Neumann, p. 14. Degraded to Argas reflexus var. magmis, 



Neumann. '1905. p. 239. 

 1929. Argas columbarum (Shaw): Oudemans, part 2, p. 138. 

 1936. Argas reflexus (Fabricius): Brumpt pp. 1187-1188. 



ADULT 



Body. — Oval, often distinctly narrower in front. Margin irregularly striate 

 with the striae varying in width in different specimens. Margin slightly turned 

 up (whence reflexus). Often larger than persicus and may reach 9.0 x 5.0 

 (Nuttall et al. 1908). Specimens may be as small as 6.25 x 3.75. 



Integument. — Integument on dorsum and venter raised into irregular, 

 sinuous wrinkles which may be short or long. Nuttall et al. (1908) state that 

 the integument is much more finely wrinkled than in persicus. Specimens from 

 the Americas show the wrinkles to be coarser. 



Discs. — Variable; may 'be either large and numerous or small and few; 

 when numerous arranged radially. 



Buttons. — Absent or few in number, often each with a short, fine hair 

 arising from the pit. 



Legs. — Moderate in length and size. Surface irregular — nearly granular. 

 Subapical dorsal protuberances well developed, about equal on all the legs. 

 Length of tarsus I (in larger specimens), 0.78; metatarsus, 0.78. Length of 

 tarsus IV, 0.96; metatarsus, 0.93. 



Coxae. — Coxae I and II separated; all others contiguous. (Nuttall et al 

 (1908) state that coxae I and II are contiguous.) 



Folds. — Coxal and supracoxal folds present; all other folds absent. 



Capitulum. — Removed from the anterior margin by about its own length 

 (including the palpi). Basis capituli nearly twice as wide as long; lateral 

 margin with a group of short, fine, erect hairs at about the middle. Entire 

 length of palpal article 1 in contact with and overlapping the base of the 

 hypostome as a knife-edge flange. Postpalpal hairs absent. 



Hypostome. — Mildly tapering; apex rounded. The larger denticles 

 arranged -/•_> with about 3 or 4 in each file. Numerous very fine denticles in 

 the middle arranged in diagonal rows (more numerous than in persicus) 



