STEUCTUKE OF LEPIDOSTROBUS. 37'i 



as Traqnairiae are the macrospores, but little can be added. An impartial examination 

 of the slides on which he based this conclusion (C.N. 1063-1071), taken in conjunction 

 with the way in which we know that vegetable structures are cruslied on and in one 

 another, leads one to the conclusion arrived at by Count Solms-Laubach, who says, 

 speaking of these same slides : •' I have seen these preparations in his collection, and T 

 must acknowledge that small groups of Traquairiae or single Traquairite do lie inside 

 many of the sporangia. But the whole specimen is so macerated that these might well 

 have found their way in from the outside while it was rotting in the water, especially as 

 other sporangia of the same cone contain small remains of tissue, which could not have 

 got there in any other way"*. 



That the sections are those of a strobilus is certain, and as in all probability it is a 

 Lepidosirohus, it is an interesting coincidence, if nothing more, to find in a slide 

 belonging to Dr. Scott (S. 83) much the same type of axis (the vascular axis alone is 

 preserved) again associated with Traquairiae. In this section, however, the diameter of 

 the vascular axis is 2 millims., whereas in the type slides of L. Traquairia it is about 

 1*2 millim. ; this increased size in the former being accompanied by an increase in the 

 magnitude of the pith, the actual thichiess of the vascular ring being the same in both 

 cases. The pith-cells (in S. 83) have the appearance of being in ^an actively meristematic 

 condition. 



On PI. XXXVII. fig. 22 I have given a diagrammatic representation of part of the cone 

 of the Lepidosirohus oldhamius type, as seen in radial longitudinal section and magnified 

 about four times. It differs from the restoration of the same cone given by Williamson f 

 in some details, as will be seen by comparison of the two figures. The longitudinal 

 course of the leaf-trace bundles in the axis is shown, as well as the origin of the parichnos 

 space, Pa.^ from the middle cortex ; the position of the ligule, U , is indicated, and the 

 great upward extension of the sporophyll laminae, la. 



Lepidostrobus foliaceus, sp. nov. 



As already mentioned, this form was figured by Williamson ('' Organization," &c.. 

 Part XIX. fig. 57, Phil. Trans. B. 1893), but the sections of this type at present known 

 are insufficient to enable me to give anything like a complete account of the strobilus. 

 However, there is no doubt as to its specific distinction from forms of the L. oldhamius 

 type. 



Referring to Williamson's figure, it will be seen that the specimen is incomplete 

 below, while the apex is well preserved. The cone is small, very much smaller 

 than any of the other forms described, its diameter being only about 1 centim , while 

 that of L. oldhamius is between 3 and 4 centims. The length of the specimen as 

 preserved is about 2"5 centims. Unfortunately the section is not quite radial, and 

 so the axis is scarcely reached ; but the thin upwardly-directed pedicels are well shown, 

 each bearing on its upper surface a single radially-elongated sporangium. 



* Solms-Laubach, 'Fossil Botany ' (English translation, 1891), pp. 184-185. 

 t " Organization," Part XIX., PhU. Trans. J5. 189^, pi. ^i. fig. 62. 



6EC0ND SEEIES. — BOTANY, VOL. V. 3 L 



