Pteridophyten. — Floristik etc. 275 



pinna-trace originales, exclusively marginally; an extra-marginal 

 origin of the pinna-trace obtains throughout the Osmundaceae, the 

 Gleicheniaceae , the Hymenophyllaceae and the Aspidiae. In the follo- 

 wing groups both modes of eniission are represented: Cyatheaceae, 

 Woodsiae, Davalliae, Aspleniae and Pterideae. The author believes 

 that the marginal mode of departure of pinna-traces is prevalent 

 in the most primitive as well as in the most highly specialized 

 forms; the extra-marginal mode of origin he believes to represent 

 an intermediate phase in the histor}^ of the supply of pinna-traces. 

 On his interpretation of the phenomena occurring in the Zygopterideae 

 this fossil family included forms with marginal and forms with 

 extra-marginal origin of pinna-traces, though he admits that the 

 Organization of the petioles of these fossils is so different from that 

 of recertt ferns that they must not be examined too criticalh^ (sie) 

 for protot5^pes of living forms. He believes that there is no funda- 

 mental difference between marginal and extra-marginal modes of 

 pinna-trace Omission, the traces for the pinnae usually going off 

 from the part of the petiolar bündle nearest to the pinnae themselves. 

 An extra-marginal mode of departure seems to have come into being 

 with the development of more or less elaborate adaxial hooks in the 

 leaf-traces, the tunction of these hooks being presumably to carry 

 forward the water-supply for the Upper part of the leaf. The origin 

 of the extra-marginal emission of pinna-traces w^as thus probably 

 caused b}^ increase in size and heavier pinnation of the frond. It is 

 believed that in some cases in which the carr3ang forward of the 

 water-supply for the upper part of the leaf is assured by abaxial 

 complications of the adaxial Strands of the petiole (e g. in Polypodiian 

 and species of Pteris) the marginal method of origin of pinna-traces 

 has been retained by advanced forms. Then after the increase in 

 size and heavier pinnation, there seems sometimes to have been a 

 "condensation" (sie) of the leaf-trace, perhaps owing to reduction in 

 size ot the leaf, making itself manifest in the breaking up of the 

 originally solid leaf-trace into separate Strands and then the adaxial 

 hooks may gradualh'- have disappeared, so that the origin of the 

 traces of the pinnae maj'^ have become once more marginal and the 

 carrying forward of the water-supply to the apex of the leaf may 

 be undertaken by the abaxial part of the trace. 



The Marattiaceae and Hehninthostachys have a peculiar origin 

 of the pinnatraces. In the specimens examined the petiolar bundles 

 were arranged in two closed circles. The author interprets this trace 

 as a modification of a horse-shoe-shaped type with much infolded 

 margins, the latter being represented by the internal Strands; as 

 both circles contribute to the vascular supply of the pinna the 

 departure of the trace of the latter cannot be described as either 

 marginal or extra-marginal. Both marginal and extra-marginal origin 

 of the pinna traces occur within the limits of the genus Botrychium. 



Isabel M. P. Browne (London). 



Benz, R. von. Neuer Fundort der Waldsteima tey^iata (Steph.) 

 Fritsch in Kärnten. (Carinthia. IL CIV. Jahrg. [=i 24. Jahrg. 

 der Car. II). p. 52—54. Klagenfurt, 1914.) 



Gabriel Höfner entdeckte 1888 die genannte Reliktpflanze aui 

 Wiesenabhängen des Prössinggrabens am Füsse der Koralpe; 

 jetzt ist die Pflanze hier bedeutend seltener. Am Burgstallkogel 

 (bei Lavamünd, unweit des Siegelsteines) sah Verf. auch die Art, 



