Pteridophyten. 201 



into an internal and an external Strand and the former "persists 

 across the tront of the leaf-trace, which appears as a ring of Strands". 

 When the trace begins to pass out a vegetative bud is formed at 

 its base, the vascular supply of the latter arising from two of the 

 Strands of the trace. Finally the leaf-trace is cut off by a scleren- 

 chymatous intrusion and the gap in the ring of dictyosteles fiUed. 

 A second species, P. Wülinckii, Moore, showed an anatomical 

 structure like that of P. alcicorne. 



P. aethiopicuni is a much larger species, with leaves about 1^ 

 feet long; it has three kinds of leaves: foliage leaves, nest leaves 

 and fertile leaves. The rhizome has a complex stelar structure; 

 there is an outer ring of meristeles, anastomosing frequently and 

 forming a perforated dictyostele. Inside this outer ring is a large 

 number of meduUary Strands, cut off from the outer Strands or 

 arising from already existing medullary bundles. The outer meris- 

 teles are concentric, without any definitely marked protoxylem. In 

 the young plant the earliest stage seems to be a haplostele, or 

 solid Strand of xylem surrounded by phloem. Phloem and paren- 

 chyraa soon appear in the interior, but there is no solenostelic 

 stage, the Strand dividing into two and then into four meristeles 

 by sclerenchymatous intrusions. These meristeles are continually 

 anastomosing from their first formation. The first permanent medul- 

 lary bündle is cut off early from one of the meristeles. 



The anatomy of the nest and foliage leaves corresponds in 

 essentials and they probably originated from a common type of 

 leaf. The mature trace consists of an outer ring of bundles arising 

 from several of the Strands of the external axial dictyostele and an 

 inner series of bundles. some of which arise from the medullary 

 bundles of the stem while others originate from the external Strands 

 of the trace. After the departure of the trace a vascular commissure 

 is formed by accessory Strands. In the young plant the trace 

 originates from a meristele that divides into two; one of these divides 

 into two bundles, but of the resulting Strands one joins a cauline 

 meristele, so that the trace consists of two Strands. These divide 

 so as to form three bundles, there being no accessory bundles in 

 the young leaves. 



It is held that anatomically Platycerium might represent the 

 dictyostelic form of a series of which Dipteris and Matonia would 

 represent the solenostelic forms. If on further investigation charac- 

 ters other than anatomical should point in this direction the position 

 of Platycerium. will probably be found to be near to the Matonia- 

 Dipteris series. Isabel Browne. 



Morton, F., Beiträge zur Kenntnis der Pteridophy ten- 

 gattung Phyllitis. fOesterr. bot Zeitschr. LXIV. 1/2. p. 19—36. 

 5 Abb. 2 Karten. 8». 1914.) 



Der erste Teil der Arbeit handelt über die Auffindung von 

 Phyllitis hemionitis (Lag.) O. Kuntze (= Scolopendrium hemionitis 

 Sw.) im Quarnerogebiet. Verf. fand dieselbe im Juni 1912 an den 

 Nordostabstürzen der norddalmatinischen Inseln Arbe und S. Gre- 

 gorio in schattigen Felsspalten. Verf. bringt sodann eine Zusam- 

 menstellung aller bisher bekannt gewordenen Standorte, die sich 

 um das Mittelmeer, namentlich um dessen westlichen Teil gruppieren. 

 Er nimmt an, dass die jetzt so sehr zerstreuten Standorte Reste 

 einer ehedem weiteren und dichteren Verbreitung sind, und hält 



