Pteridophyten. — Fioristik, Geographie, Systematik etc. 631 



Doodia also shows a certain displacement of the separate sori arising 

 from the disintegration of the fusion-sorus; these no longer form 

 a Single row on each side of the midrib and it would seem that as 

 well as displacement there has been an Initiation of sori in positions 

 hitherto untenanted by them. 



There is evidence of suppressed pinnation in Scolopendrium 

 vulgare L. and of suppressed or imperfectl}'" developed bipinnation 

 in B. punctulatum var. Krebsii. The author believes that where the 

 suppression of pinnation is less perfect than in some species of 

 Scolopendrium the sori of the suppressed pinnae may develop as 

 extra sori between the veins. If such additional sori were extended 

 downwards beyond the point of branching of the vein that supports 

 them — a condition that actually occurs in B. punctulatum var. 

 Krebsii and in Camptosaurus — we should reach the soral distri- 

 bution characteristic of the subgenus Diplasium. of Asplenium. In 

 this subgenus the sori are back to back on unforked veins. Diplasium 

 is the subgenus of Asplenium in which it is most difificult to recognize 

 a Blechnoid affinity in the distribution of the sori. 



Professor Bower maintains that the non-soralcondition of certain 

 Acrostichoid forms is attained by the spreading of the sori, the 

 sporangia being no longer restricted to the region over the veins; 

 analogies for such a process were found in species of Blechnum, 

 e. g. B. filiforme (A. Cunn) Ettighsn. B. pennamarina (Poir) and 

 StenocJilaena sorbifolia. The author recognises several minor lines 

 in the Acrostichoid series. Like the genus Polypodium the old genus 

 Acrostichum is not a phyletic genus; the ferns included in it show 

 conditions of soral construction probably reached along several 

 distinct though related phyletic lines. 



The majority of the ferns dealt with in this paper have sori of 

 the mixed type; but gradate sori also occur especially in the simpler 

 forms such as Matteuccia intermedia and Blechnuyn tabulare. In 

 several species of Blechnum , e. g. B. (Eu-Blechnum) brasiliense the 

 sorus is at first gradate but becomes mixed. In Blechnum {Lomaria) 

 alternatum the sporangia of a sorus arise simultaneously ; in Steno- 

 chlaena sorbifolia the sori ma}'' be of the simultaneous or of the 

 mixed type. 



As in the Cyatheoid Ferns, so in the Blechnoid series Professor 

 Bower regards the creeping form of axis as more primitive than 

 the upright; he also thinks that the simple pinnation preponderant 

 though not universal in the group has been derived from bipinnation. 

 All the forms examined are, when mature, dictyostelic, though in 

 Plagiogyria the axis is not far removed from solenostely. In the 

 simpler forms the leaf trace originates as two Strands which maintain 

 their identity for some distance and then divide to form in the 

 petiole a horse-shoeshaped series of bundles; in the more complex 

 forms, e. g. Stenochlaena sorbifolia and Woodwardia radicans, the 

 trace originates as five or six bundles. Finally, in some forms 

 vascular fusions take place in the petiole. 



Isabel Browne (Londen). 



Makino, T., Observations on the flora of Japan, [cont], 

 (Bot. Mag. Tokyo. XXVIII. p. 20-30, 31—36, 105-130, 153-160, 

 165-170. to be cont. 1914.) 



The first part of this new series of papers gives a new genus 

 Physaliastrum Makino nov. genus, closely allied to Physalis ■A.ndi to 



