Jan., 1921] Translocation of Carbohydrate Materials 



91 



of temperature during the growing and dormant periods are 

 accompanied by changes in acidity in the plant, which in turn 

 affect the activity of the enzymes which have always been 

 found by actual experiment to be present at all seasons of the 

 year. 



The data of the analyses are presented under three headings : 

 First, curves showing changes in carbohydrates in fruit spurs; 

 second, curves showing changes in carbohydrates in two-year-old 

 seedlings; third, curves showing acidity in both the fruit spurs 

 and seedlings. The curves in each case are followed by a brief 

 summary and discussion. 



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Fig. 2. Curves showing the percentage of sucrose, glucose and maltose in 

 fruit spurs of apple from May to April, 1919-1920. The ordinates represent the 

 percentage of sugars and the abscissae, the season of the year. 



The Fruit Spurs. 



1. Change of temperature has a marked bearing on the 

 translocation of carbohydrate materials in fruit spurs of apple. 



2. The accumulation of both total carbohydrate and 

 starch shows marked increase during August and September, the 

 maximum accumulation occurring in September. 



3. As the temperature decreases in autumn the starch 

 content decreases without any marked increase in either the 

 total or reducing sugars. This decrease of starch continues 

 until February, when it reaches its maximum for the dormant 

 season. The total carbohydrate content of the fruit spurs 

 follows a similar course. 



