SMALL 

 IMPORT 



LOW STORAGE 

 LOW BIOMASS 

 LOW PRODUCTIVITY 



SMALL 

 EXPORT 



LARGE 

 IMPORT 



MODERATE 

 EXPORT 



HIGH STORAGE 



HIGH BIOMASS 



HIGH PRODUCTIVITY 



Figure 7. The hypothetical relationship between nutrient input (excluding carbon), 

 biomass, primary productivity, and nutrient export (including carbon) from mangrove 

 ecosystems. Top: small nutrient import. Bottom: large nutrient import. 



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