BY R. J. TILLYARD. 



649 



position. In the hindwing, the whole anal area is devoid of 

 veins, except for one very close to the wing-border, the homo- 

 logies of which cannot be determined with certainty, and a vein 

 running below Cu^ in the usual position of Cu 2 , which may be 

 either that vein or 1A. 



In the forewing, M is very completely fused basally with Cu, 

 so that all traces of the cubito-median Y-vein are lost. The 

 main stem of M 1 _ 4 is obsolescent as far as mf. In the hind- 

 wing, this main stem is more clearly shown, and is not fused 

 with Cu; a tiny remnant of the cubito-median Y-vein appears 

 to be visible at the extreme base of the wing. 



The discovery of this family is chiefly interesting in that it 

 proves the existence of hitherto unsuspected Homoneurous types. 

 It may reasonably be hoped that further discoveries along simi- 

 lar lines may yet be made. Indeed, the discovery of the re- 

 markable genus Anomoses in South Queensland by Dr. A. J. 

 Turner (31) is probably a case in point; for this genus does 

 not appear to be a true Jugo-frenate type, and should pro- 

 bably be placed either in the Prototheoridue or in a new family 

 Anomosetidae. 



In concluding this account of the Suborder Homoneura, it is 

 interesting to compare the wing-venational characters of the 

 five families represented therein, in such a way that the archaic 

 characters in each family may be readily picked out. This is 

 best done by means of a Table (Table iii.), in which the most 

 archaic condition of each separate character is represented 

 under the letter A, while B, C, . . . represent more spe- 

 cialised conditions of the same, in ascending order. The Arche- 

 type of the Homoneura w 7 ill possess all the archaic conditions 

 indicated under A. The "percentage archaism" of that Arche- 

 type will therefore be 100. For comparison with this, the 

 "percentage archaism" of each of the five families is worked 

 out in the last column of the Table. When w t <; have studied the 

 Heteroneura, and tabulated their characters in the same man- 

 ner, it will be seen that that Suborder does not possess a single 

 archaic condition which is not also to be found sornewnere 

 within the Suborder Homoneura/ In other words, the Arche- 

 type of the Homoneura is also the Archetype of the Order 

 Lepidoptera as a whole. 



