070 



THE PANORPOID COMPLEX, 111., 



areoeel between R g and R4 + 5 (in Wingia), or, in those cases 

 where the latter is not stalked beyond the areoeel, between Rs 

 and R 4 (as in Carpoeapsa) . Unless this cross-vein had been 

 present, to furnish the channel along which the new trachea 

 grew out from R 3> no areoeel could have been formed. Thus 

 we are bound to conclude that the cross-vein ir was present in 

 the ancestral forms of the Tortricina and Tineina.* 



Text-Fig.92. 

 Details of tracheation of the imaginal forewing, freshly emerged, of 

 Wingia lambertiella (Wing), (fam. Oecophoridae) . (x 11). Note 

 the shrivelled median trachea (Alt) and cubital trachea (Cut). Let- 

 tering as on p. 535. 



Another striking change at metamorphosis in Wingia 

 is the complete and quite sudden switching off of the 

 trachea Ciij near its point of origin on' Cu, across 



to M, via M 



5- 



In the latest pupal wings which 



I examined, I could find no trace of a trachea in M 5) though I 

 looked carefully for it. But, in the fresh imaginal wing, a 

 strong trachea proceeds from the base of M, curving down 

 along Mi 5 (which, in this moth, is not at all well aligned with 

 Cui), and supplying Ciij right to its ends. Above this trachea 

 could be seen the shrivelled remnant of trachea M 1 _ 4) while two 

 much finer trachea?, proceeding from R, more or less followed the 

 line of M distad from it (Text-fig. 92) . Below M 5 , the rem- 

 nant of trachea Cu was plainly visible, with its original dich- 

 otomy into C\\ and Cu 2 , the former still exactly in line with 



* Further proof of this fact is to be found in the existence of a number 

 of genera, in both these superfamilies, in which the areole or radial cell 

 is retained in its entirety, and in every such case this cell is closed 

 by ir. (See Turner, 32, figs.24-30). 



