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Comments 



Auerbach: It seems to me that the examples discussed under I and II 

 are not at all equivalent. In I, the equilibrium refers to a pair of alleles, 

 in II to whole genomes, as in Oenothera, or whole chromosomes, as in 

 Dobzhansky's Drosophila inversions. The balance in Oenothera is upheld 

 by genes which are not allelic. This difference between equilibrium 

 for alleles or for larger parts of the genome may not be important for 

 the way a population achieves equilibrium in nature, but it is the 

 important question in the assessment of genetical radiation damage. As 

 far as I know, there are few examples for selectional equilibrium at 

 the allele-level, the only level which has to be considered in this regard. 



