EVIDENCES FROM PALAEONTOLOGY 147 



Were there time enough to do so, we might trace the development 

 of this family backward, step by step, through all the many stages 

 between the Pleistocene and the Upper Eocene in quite as unbroken 

 sequence and in as full detail as can be done for the horses. We must, 

 however, pass over all the intermediate steps and consider the ances- 

 tral camels of the Upper Eocene. These were very little animals, 

 hardly larger than a jack rabbit, which had the full complement of 

 teeth, 44 in total number, and all with very low crowns. The Umbs, 

 and especially the feet, are relatively short, the ulna is complete and 

 separate, as is also the fibula; there are four toes m each foot, though 

 the lateral pair of the hind foot are extremely slender, and there is no 

 co-ossification to form cannon-bones. The hoofs are well developed, 

 in form like those of an antelope, so that there can have been no pad. 

 For the present, the line cannot be carried back of the Upper Eocene, 

 the probable ancestors from the middle and Lower Eocene being, as 

 yet, represented only by fragmentary specimens. 



In addition to this main stem of cameline descent which resulted 

 in the modem species, there were two short-lived side branches which 

 should be mentioned. One, ending in the Lower Miocene, was the 

 series descriptively called "gazelle-camels," small animals with very 

 long and slender legs, evidently swift runners. The other series, the 

 so-called "giraffe-camels," terminated in the Upper Miocene; these 

 were browsers and display an increasing stature, especially in the 

 length of the neck and fore Umbs. They adapted themselves to the 

 growing aridity of the western plains. 



EVOLUTION OF THE ELEPHANTS* 

 A. FRANKLIN SHTJLL 



The mastodon-elephant series shows a larger number of obvious 

 changes than most of the other series named, all of these changes 

 except that of the body having to do with features of the head. 

 From the numerous specimens of elephant-like forms available, the 

 following are selected (following Lull) as probably representing a 

 direct line of evolution: Moeritherium from the Upper Eocene of 

 Egypt; Palaeomastodon from the Lower Oliogocene of Egypt, also 

 from Lidia; Trilophodon from the Miocene of Europe, Africa, and 

 North America; Mastodon from the Pliocene and Pleistocene of 



• From A. F.' ShuU, Principles of Atiimal Biology (copyright 1920). Used by 

 special permission of the publishers, The McGraw-Hill Book Company. 



