•ECOSYSTEM BOUNDARY- 



ATMOSPHERIC 

 COMPARTMENT 



Wind-blown particulolas 



end gosftft abovs and 



bslow ground 



Biological uplak* 



of go»et 

 Biological rel9os« of 



gases, orgomc aerosols, 

 evopotronspirolion 



-INTRASYSTEM CYCLE 



Meteorologic 

 Geologic 

 Biologic 



BIOSPHERE 



Figure 9-5. Major storage compartments and flows of nutrients within the 

 forest system (from Bormann and Likens 1979). 



9-28 



