Eisen, The Chromoplasts and the Chromioles. 



133 



It must suffice to state here that in time the leaders separate 

 from each other, but to each leader, which now may be termed a 

 chromosome, remains attached a part of the original chromoplast. 

 It may naturally be asked how do we know that such is the case. 

 The answer is this: The chromoplast can nearly always be recognized 

 by its structure. It always contains certain highly refractive granules, 

 which are seen in the cbromoplasts no matter in what stage of deve- 

 lopment the nucleus may be. These granules are the landmarks by 

 which the chrouioplasts may always be recognized. Thus, when the 

 chromosomes are in the equator of the cell, the chromoplast is seen 

 in one side of the centre in the ring- like chromosome. The subsequent 

 division of the chromosome is through a simple equation, and passing 

 through the chromoplast, gives each new or daughter-chromosome an 

 equal share of the chromoplast. The chromoplast is now situated at 

 the end of one of the limbs of the horse- shoe -shaped chromosome. 

 The next stage is an umbrella or ring -like nucleus. This stage, far 

 from being caused by imperfect fixing methods, is an absolute neces- 

 sity, as being the only means by which the chrouioplasts can change 

 place. Let us understand this clearly. Before the chromosomes enter 

 the ring- like nucleus the chromoplast is found at the very point of 

 one of the limbs. When in the next cell the spermatocyte - - the 

 chromosomes reappear, we find that the chromoplast has changed its 

 position. It is no longer found at the apex of one of the limbs, but 

 is seen to be located at the very angle where the two limbs meet. 

 This change of locality takes place in the ring-shaped nucleus, in 

 which the chromomeres and chromosomes loose their individuality, the 

 chromioles and chrornoplasts remaining. 



l. 



lininoplast or true 

 nucleolus 



nucleus 

 chromoplast 



cell-wall 

 chromoplast 

 or net-knot 



1. Polymorphous spermatogoniutu. The black, dark granules are the chroinioles. 



The light colored granules are the linin granules, here represented as a thread 



only. The cell is in perfect rest. Only a few chromioles are indicated. 



