134 



Eisen, The Chromoplasts and the Chromioles. 



I liuve found the chromioles and the chromoplasts iu all cells 

 capable of mitosis which I have investigated, and I have come to the 

 conclusion that they are constant elements in all cells. The leucocyte 

 is of almost identical structure with the polymorphous spermatogonium. 

 We find in it the scattered chromioles, and among- them a varying 

 number of chromoplasts, the latter of larger size and more intense 

 staining- quality. 



The liuinoplast, or true nucleolus, is, according- to my conclusions, 

 principally a storage reservoir of linin granules, and has somewhat the 

 same relative influence on the linin granules as the chromoplasts have 

 on the chromioles. 



chromioles supported by linin 



chromoplasts 



lininoplasts 



nucleus 



leaders 



2. Polymorphous sperinatogouiuin. Two chromoplasts connected with 12 leaders 

 or incipient chromosomes. Each leader with chromomeres containing 3 chro- 

 mioles each. Three lininoplasts. 



Again, regarding the chromioles: 



These minute bodies are the only constant parts of the chromo- 

 somes. The formation of the chromosomes is due to an independent 

 process, so to say, presided over by the chrornoplast, and in every 

 respect independent of the centrosomes. The mitosis as a whole is 

 the result of two parallel but independent processes, which run parallel 

 for a time, but touch at certain points and aid each other in accom- 

 plishing their mutual ultimate purpose. In other words, the chromo- 

 plasts prepare and form the chromosomes, the centrosomes accomplish 

 their proper mitosis. 



The number of the chromioles is constant in every species. In 

 Batrachoseps the perfect chromosome contains six chromomeres, three 

 iu each limb of the chromosome. Each chromomere contains six chro- 

 mioles, which makes for each chromosome 30 chromioles, and for the 

 whole nucleus 432 chromioles. 



