coral 

 "read 

 coral 

 1 ocat 

 Sw 

 coral 

 the 1 

 i n c 1 u 

 1977; 

 water 

 predi 

 Lang, 

 prese 

 appen 

 cl ose 

 and " 

 (Lang 

 and d 

 compe 

 (Ell i 



, sw 



y" t 



s wh 

 i on 

 eepe 

 Mon 



eepe 

 o pa 

 i ch 

 on t 

 r te 

 tast 



r te 

 rt i c 

 g row 

 he c 

 ntac 

 raea 



ntacl 

 i pate 



too 

 ol ony 

 1 es c 



cave 



es a 



i n 

 cl OS 



rel 

 ommo 

 rnos 



ocat 

 de: 



Ri c 



cur 

 ctab 



unp 

 nt , 

 dage 



gro 

 poly 

 , 19 

 eve! 

 titi 

 s an 



i on 



con 

 ha rd 

 rent 

 1 e p 

 ub . 

 swee 

 s (L 

 wth 

 f unc 

 79). 

 opme 

 ve i 

 d So 



of s 

 cent 

 son , 



s (P 

 atte 

 data 

 pers 

 ewi s 

 of a 

 ti on 

 He 

 nt o 

 nter 

 1 and 



weepe 

 ratio 

 et a^ 

 rice, 

 rn of 



). P 

 on M 

 and 

 dj ace 

 al" s 

 re we 

 f swe 

 actio 

 er) . 



r te 

 n a r 



1-. 



197 



di s 

 erha 

 . ca 



re r 

 comp 

 e (d 

 at i v 

 nly 

 a (L 

 ntac 

 ound 

 1979 

 3, 1 

 trib 

 PS i 

 vern 



out i 

 etit 

 epen 

 e to 

 a re 

 i nna 

 1 es 

 col 

 ) ; m 

 n de 

 ut i o 

 n pa 

 osa 



Pric 

 nt c 

 true 

 pre 

 eper 

 ns w 



e, 1 



ompe 

 ture 

 sent 

 ten 

 i th 



975) 

 ti to 

 s wh 

 pre 

 tacl 

 the 



nely 

 i ve i 

 ding, 

 that 

 seen 

 eus ) . 

 on M. 

 ony p 

 ax ima 

 n Har 

 n ove 

 rt be 

 have 

 , def 

 rs (R 

 i ch m 

 1 imi n 

 es on 

 conge 



pres 

 nter 



of 



of 

 on c 

 De 



cav 



ent an 

 action 

 course 

 nea rby 

 ol on i e 

 scri pt 

 ernosa 



enm 

 1 ex 

 tog, 

 r co 

 caus 

 been 

 ens i 

 i cha 

 ight 

 ary 

 M. 

 neri 



eters 

 pansi o 



1977 ) 

 1 ony s 

 e they 



desc r 

 ve str 

 rdson , 



serve 

 data o 

 cavern 



d a r 

 s wi 

 , on 



com 

 s of 

 i ve 



var 

 ( den 

 n i n 

 ; or 

 urfa 



are 

 i bed 

 uctu 



et 



bot 

 n th 

 osa 



c speci es 



e therefore 

 th neighboring 



their 

 pet i tors ) . 



the Caribbean 

 patterns of 

 i ousl y 



Hartog , 



response to 



a less 

 ces ( J .C . 



usually 



as f eedi ng 

 res to deter 

 al., 1979), 

 h functions 

 e behavior 

 in artificial 

 M . annularis 



MATERIALS AND METHODS 



Th 

 River 

 dive 

 Col on 

 c i n d e 

 for m 

 rem a i 

 when 



Ou 

 other 

 expan 

 col on 

 on th 

 night 

 annul 



ese e 

 Cany 



in t h 

 i es o 

 r bio 

 ethod 

 ned b 

 their 

 r i ni 



cora 

 si on 

 i es o 

 e M. 



afte 

 a ri s 



conta 

 M. ca 

 resul 

 i m p o s 

 du ri n 

 Ne 

 the 1 

 M. £a^ 

 al rea 

 obser 

 each 

 col on 

 both 

 caver 



ct wi 

 verno 



ting 

 s i b 1 e 

 g the 

 vert h 

 onger 

 ve rno 



dy we 

 ved o 

 obser 

 i es w 

 coral 

 nosa 



to the M. 



x pen m 

 on , St 

 e NOAA 

 f M. a_ 

 ck s us 

 s) . C 

 etween 

 polyp 

 t i a 1 i 

 Is cl o 

 patter 

 f M. a_ 

 cavern 

 r cora 

 digest 

 th the 

 sa rem 

 wound s 

 to ob 

 rem a i 

 el ess , 

 -term 

 sa on 

 1 1 doc 

 n six 

 v a t i o n 

 ere ca 

 s were 

 wa s d i 

 a nnul a 



ents 

 . Cro 

 NULS 

 nnul a 



were c 



i x . T 



II Un 



ri s an 



l ng u 

 oral s 

 pair 

 s and 

 ntent 

 se to 

 ns of 

 nnul a 



nderwa 

 were 



ed col 

 t i s s u 

 was t 

 col on 

 its s 



ri s , 1 



osa c 

 1 s we 

 ed ne 

 i r t i 

 a i ned 

 . Th 

 serve 

 nder 



we w 

 conse 

 which 

 ument 

 night 

 , the 

 ref u 1 



reco 

 v i d e d 

 ri s . 



1 o n i e 

 re cem 

 a rby e 

 ssues. 



cont r 

 is unp 



the b 

 of the 

 ere pr 

 quence 



the 1 

 ed. T 

 s over 



posi t 



1 y map 

 rded . 



into 

 Bound 



ondu 

 hey 

 derw 

 d M. 

 ter 

 arra 

 on i e 

 es w 

 o ex 

 i es 

 weep 

 ocat 

 s we 

 ente 

 xpa n 

 Su 

 acte 

 redi 

 ehav 



sat 

 ov i d 

 s of 

 ocat 

 hese 



the 

 i ons 

 ped 



For 

 regi 

 ari e 



cted 

 we re 

 ater 

 cave 



at a de 

 i n i t i a t 

 Habitat 

 rnosa ( 



epoxy 

 nged 

 s of 

 e re c 

 amine 

 of M. 

 ers. 

 i ons 

 re ca 

 d i n t 

 ded p 

 bsequ 

 d i n 

 cted 

 i or o 

 urat i 

 ed wi 

 such 

 i on o 

 i nte 

 foil 

 of s 

 and t 

 data 

 ons a 

 s of 



-putty 

 so that 

 M . annu 



pth 

 ed d 



i n 

 N=8) 

 ( see 



a g 

 1 a ri 



ont ract 

 whet he 

 cavern 



ed d 

 r i n 

 osa 



Thus , 

 of e x i s 

 ref ul 1 y 

 o place 

 olyps o 

 ently , 

 the are 

 b e h a v i o 

 f their 

 on dive 

 th the 



i ntera 

 f sweep 

 ract i on 

 owing t 

 weepers 

 he beha 



analys 

 d j a c e n t 

 these " 



pri o 

 ting 



map 

 , al 

 f M. 

 poly 

 a su 

 ral 



swe 



of 60 

 u r i ng 

 March 



were 



Chorn 

 ap of 

 sand 

 uri ng 

 troduc 

 woul d 

 r to i 



sweep 

 ped. 

 1 col o 



caver 



ps of 

 rround 

 respon 

 eper t 



feet 

 a sat 

 of 19 

 cemen 

 esky, 

 about 

 M . c_a 

 the d 

 ti on 

 af fee 

 nt rod 

 er te 

 The f 

 ni es 

 nosa 

 the d 

 i ng t 

 se ma 

 en tac 



i n Sal t 

 urati on 

 82. 

 ted to 



1983, 



1 cm 

 vernosa 



ay. 

 of 



t the 

 uc i ng 

 ntacl es 

 i rst 

 Of M. 

 i n 



i g e s t e d 

 he 



de it 

 1 es 



opportunity to follow 

 ctions for colonies of 

 er tentacles wa s 

 s subsequentl y we re 

 wo months. During 



on the M. cavernosa 

 vior and condition of 

 is, each col ony of M. 



to and not adjacent 

 adjacent" and "non- 



62 



